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Sceloporus dixoni BRYSON & GRUMMER, 2021

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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymSceloporus dixoni BRYSON & GRUMMER 2021
Sceloporus aeneus – DUELLMAN 1965 (in part)
Sceloporus aeneus – THOMAS & DIXON 1976 (in part)
Sceloporus aeneus – BENABIB et al. 1997 (in part)
Sceloporus aeneus aeneus – SMITH 1937 (in part)
Sceloporus aeneus aeneus – SMITH 1939 (in part)
Sceloporus aeneus aeneus – SCHMIDT & SHANNON 1947
Sceloporus aeneus aeneus – DUELLMAN 1961 (in part)
Sceloporus aeneus aeneus – MINK et al. 1996 (in part)
Sceloporus aeneus subniger – SMITH et al. 1993 (in part)
Sceloporus aeneus subniger – BRYSON et al. 2012 (in part)
Sceloporus subniger – GRUMMER et al. 2014 (in part)
Sceloporus dixoni — HEIMES 2022 
DistributionMexico (Jalisco)

Type locality: Nevado de Colima, 13.5 mi W Cd. Guzmán, municipality of San Gabriel, Jalisco (N 19.6427°, W 103.6236°, 2375 m; WGS84);  
Reproduction 
TypesHolotype: UTA 61714, adult male, (field number RWB 0649), collected 24 June 2006 by R. W. Bryson Jr.
Paratypes: MZFC 22053, 22054; UTA 61713, 61715–61716). Michoacán: 11.7 mi W Zacapu on rd to Zamora (MZFC 22055, 22056; UTA 61699–61702). 22 km N Uruapan on Hwy 37 (UTA 61703, 61704), same data as holotype. 
DiagnosisDiagnosis: Sceloporus dixoni is a member of the S. scalaris group, sharing with other species in this group parallel lateral scale rows, femoral pore series that are either in contact or separated by no more than two scales, females with smooth preanal scales, and males with lateral abdominal color patches (Smith 1939; Smith et al. 1997; Watkins-Cowell et al. 2006). Sceloporus dixoni can be distinguished from other species in this group by the following combination of characters: single canthal on each side of the head, small adult size (maximum SVL = 54 mm, average 47.1 mm), 37–45 dorsal scales (average 41), 37–43 scales around midbody (average 40), 32–39 ventral scales (average 35), tibia length/head length proportion of 0.76–0.94 (average 0.86), 4–5 supralabial scales (mode of 5), 12–18 scales bordering the interpariatel scale (average 15), 31–36 femoral pores in males (average 34), adult females with lightly mottled venters, and adult males with extensive dark pigment on the venter, heavily mottled throats, and orange or rust-colored flanks (BRYSON & GRUMMER 2021).


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CommentFor additional references see Bryson & Grummer 2021 (not provided upon request)

Synonymy: Thomas & Dixon (1976) argued that S. a. aeneus and S. s. subniger were indistinguishable. Smith et al. (1993) challenged this conclusion, claiming it was based on misidentified specimens from Nevado de Toluca and therefore an inaccurate description of the status and distribution of S. a. subniger. Sceloporus subniger and S. aeneus were subsequently considered distinct species in checklists (Liner 1994; Bell et al. 2003)

Distribution: see Bryson & Grummer 2021: 342 (Fig. 2) for a (rough) map as S. subniger “West” (not dixoni). 
EtymologyThe specific epithet is a patronym honoring the late James R. Dixon for his decades of research on Mexican herpetofauna, including several insightful studies of the S. scalaris group. “Doc” Dixon took an early interest in the academic growth of the first author and made a profound and lasting impact. For this and for his encouragement and support, he will be truly missed. 
References
  • BRYSON, ROBERT W.; JR., JARED A. GRUMMER, ELIZABETH M. CONNORS, JOSEPH TIRPAK, JOHN E. MCCORMACK, JOHN KLICKA 2021. Cryptic diversity across the Trans-Mexican Volcanic Belt of Mexico in the montane bunchgrass lizard Sceloporus subniger (Squamata: Phrynosomatidae). Zootaxa 4963 (2): 335–353 - get paper here
  • Heimes, P. 2022. LIZARDS OF MEXICO - Part 1 Iguanian lizards. Edition Chimaira, Frankfurt Am Main, 448 pp. - get paper here
 
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