You are here » home › advanced search › search results › Sceloporus grammicus

Sceloporus grammicus WIEGMANN, 1828

IUCN Red List - Sceloporus grammicus - Least Concern, LC
iNaturalist logo

Can you confirm these amateur observations of Sceloporus grammicus?

Add your own observation of
Sceloporus grammicus »

Find more photos by Google images search: Google images

Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
SubspeciesSceloporus grammicus disparilis STEJNEGER 1916
Sceloporus grammicus grammicus WIEGMANN 1828
Sceloporus grammicus microlepidotus WIEGMANN 1834
Sceloporus grammicus tamaulipensis SITES & DIXON 1981 
Common NamesE: Graphic Spiny Lizard
S: Lagartija de Arbol del Sur
E: (Northern) Mesquite Lizard, Northern Plateau Graphic Lizard [microlepidotus]
S: Lagarto de Mesquite [microlepidotus]
E: Mezquite Graphic Lizard [disparilis]
S: Lagartija de Arbol del Noreste [disparilis]
E: Tamaulipas Graphic Lizard, Sierra de Tamaulipas Graphic Lizard [tamaulipensis]
S: Lagartija de Arbol de Tamaulipas [tamaulipensis] 
SynonymSceloporus grammicus WIEGMANN 1828: 370
Sceloporus pleurostictus WIEGMANN 1828: 370
Tropidurus grammicus — WAGLER 1830
Tropidolepis pleurostictus — GRAY in CUVIER 1831
Tropidolepis grammicus — GRAY in CUVIER 1831
Tropidolepis grammicus — DUMÉRIL & BIBRON 1837: 306
Tropidolepis Grammicus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 270
Sceloporus grammicus — BOULENGER 1885: 231
Sceloporus grammicus — SMITH 1939: 179
Sceloporus grammicus grammicus — SMITH & TAYLOR 1950: 119
Sceloporus grammicus — LINER 1994
Sceloporus grammicus — LARA-GÓNGORO 2004: 25
Sceloporus gramicus — OCHOA-OCHOA & FLORES-VILLELA 2006 (in error)
Sceloporus grammicus — WIENS et a. 2010
Sceloporus grammicus — SÁNCHEZ-RIVERA et al. 2022
Sceloporus grammicus — HEIMES 2022

Sceloporus grammicus disparilis STEJNEGER 1916
Sceloporus disparilis STEJNEGER 1916: 227
Sceloporus microlepidotus disparilis — DUNN 1936: 472
Sceloporus microlepidotus microlepidotus — DUNN 1936: 472
Sceloporus pilsbryi DUNN 1936: 473
Sceloporus microlepidotus disparilis — GLOYD 1942
Sceloporus microlepidotus disparilis — SCHMIDT & OWENS 1944
Sceloporus grammicus disparilis — SMITH & LAUFE 1945: 332
Sceloporus grammicus disparilis — SMITH 1946
Sceloporus grammicus disparilis — SMITH & TAYLOR 1950: 120
Sceloporus grammicus disparilis — TAYLOR 1952
Sceloporus grammicus disparilis — WILLIAMS et al. 1960
Sceloporus grammicus disparilis — VAN DEVENDER & LOWE 1977
Sceloporus grammicus disparilis — LARA-GONGORA 1986
Sceloporus grammicus disparilis — TANNER 1987
Sceloporus disparilis — BELL et al. 2003
Sceloporus disparilis — LARA-GÓNGORO 2004: 25

Sceloporus grammicus microlepidotus WIEGMANN 1834
Sceloporus microlepidotus WIEGMANN 1834: 370
Tropidolepis microlepidotus — DUMÉRIL & BIBRON 1837: 308
Tropidolepis Microlepidotus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 270
Sceloporus microlepidotus — BOULENGER 1885: 232
Sceloporus rubriventris GÜNTHER 1890 (fide BELL et al. 2003)
Sceloporus microlepidotus microlepidotus — SMITH 1939: 184 (part.)
Sceloporus grammicus microlepidotus — SMITH & LAUFE 1945: 332
Sceloporus dispar BAIRD & GIRARD 1852: 127 (fide SMITH & LAUFE 1945)
Sceloporus heterurus COPE 1867: 322
Sceloporus microlepidopterus — HERRERA 1890: 331 (error typographicus)
Sceloporus microlepis — BOULENGER 1894: 731 (error typographicus)
Sceloporus microlepidotus — BOULENGER 1897
Sceloporus microlepidotus microlepidotus — DUNN 1936: 472
Sceloporus microlepidotus microlepidotus — SCHMIDT 1947
Sceloporus grammicus microlepidotus — SMITH & TAYLOR 1950: 120
Sceloporus grammicus microlepidotus — CONANT & COLLINS 1991: 104
Sceloporus grammicus microlepidotus — BELL et al. 2003
Sceloporus grammicus microlepidotus — CASTRO-FRANCO & BUSTOS-ZAGAL 2003
Sceloporus microlepidotus — LARA-GÓNGORO 2004: 25
Sceloporus microlepidotus — ARÉCHAGA-OCAMPO et al. 2008
Sceloporus grammicus microlepidotus — CROTHER et al. 2012
Sceloporus microlepidotus — ROTH-MONZÓN et al. 2018
Sceloporus grammicus microlepidotus — ILLESCAS-APARICIO et al. 2016

Sceloporus grammicus tamaulipensis SITES & DIXON 1981
Sceloporus grammicus tamaulipensis — BELL et al. 2003
Sceloporus tamaulipensis — LARA-GÓNGORO 2004: 25 
DistributionUSA (S Texas),
Mexico (Chihuahua, Durango, Zacatecas, Coahuila, San Luis Potosi, Nuevo Leon, Tamaulipas, Oaxaca, Guerrero, Quéretaro, Aguascalientes, Nayarit, Colima)

microlepidotus (partly fide SMITH & TAYLOR 1950): Mexico (Jalisco, N Guanajuato, N Hidalgo, Colima, Distrito Federal, Oaxaca, Puebla, Veracruz, México, Morelos, Tlaxcala, Michoacán), USA (Texas); Type locality: Mexico. Restricted to México, Distrito Federal, by SMITH & TAYLOR 1950.

disparilis (fide SMITH & TAYLOR 1950): Mexico (N Hidalgo, Guanajuato, S Zacatecas, Chihuahua, Coahuila, Durango, Nuevo Leon, San Luis Potosi, Tamaulipas), USA (S Texas). Type locality: Lomita Ranch, 6 miles north of Hidalgo, Texas.

heterurus: Type Locality: Mirador, Veracruz, Mexico.

pilsbryi: Type Locality: Alvarez, 7500 ft (elevation), San Luis Potosí, Mexico.

pleurostictus: Type locality: "Mexico", restricted to Chilpancingo, Guerrero, Mexico, by Smith and Taylor (1950).

rubriventris: Type Locality: Omilteme {= Omiltemi), Guerrero, Mexico.

Type Locality: In oak forest 4.3 rd km S Hacienda Acuña, and 45.3 rd km N Gonzales in the Sierra de Tamaulipas, in southern Tamaulipas, Mexico.

tamaulipensis: Type locality: Mexico. Restricted to Chilpancingo, Guerrero, by SMITH & TAYLOR 1950.  
Reproductionviviparous (ovoviviparous); all species of the grammicus group are ovovivparous. 
TypesSyntypes: ZMB 641-643, collected by F. Deppe.
Holotype: USNM 33041, collected by William Lloyd, 17 June 1891 [disparilis]
Holotype: USNM 25064, Probably, given incorrectly in original description as USNM 6589, collected by Charles Sartorius (Cochran, 1961) [heterurus]
Holotype: ANSP 20085, collected in 1934 by Henry A. Pilsbry, Frances W. Pennell and Cyril H. Harvey [pilsbryi]
Type: ZMB (Presumably, but not listed in Taylor, 1969), Ferdinand Deppe collector [pleurostictus]
Syntypes: BMNH 1946.8.10.23-24, formerly 1889.11.13.54-55, collected by F. D. Godman [rubriventris]
Holotype: TCWC 57386, collected 23 July 1979 by J. W. Sites, Jr., and J. R. Dixon [tamaulipensis]
Holotype: ZMB [microlepidotus] 
DiagnosisDiagnosis: Smith 1939: 179


Additional details (2719 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy: Sceloporus grammicus disparilis has been synonymized with Sceloporus grammicus microlepidotus by SITES & DIXON 1981. This species shows extreme karyotypic variation, both inter- and intrapopulational (Sites 1983). Synonymy partly after SMITH & TAYLOR 1950, who also mention intergrades between grammicus grammicus and grammicus microlepidotus from Cerro Sàn Felipe and Cerro San Juan north of Oaxaca City. Sceloporus rubriventris GÜNTHER 1890 has been placed in the synonymy of S. grammicus by SMITH & TAYLOR 1950.

Because of the isolation of the tamaulipensis population BELL et al. (2003) suggested that it should be regarded as a species. 
EtymologyNamed after the Greek adjective gramma, meaning script, or graphic, referring to the lines that cross the gray back. This is one of the most characteristic features of this group.

dispar: The Latin noun dispar means "unlike," or "different", referring to the dorsal scales, which were much smaller than the tail scales.

disparilis: The Latin dispar "unlike," or "different," and -ilis, "having the quality of," refer to the distinctive color and nature of this species.

microlepidotus: The Greek micro-, "small," and lepidos, "scale," refer to the small dorsal body scales.

pilsbryi: This Latin noun in the genitive singular honors Henry A. Pilsbry, an outstanding conchologist at the Academy of Natural Sciences of Philadelphia in the late 19th and early 20th centuries.

pleurostictus: The Greek pleura, "side," and stiktos, "spotted" refer to the scattered dark spots on sides of body.

rubriventris: The Latin nouns rubra, "red," venter, "belly," and the Greek suffix -tris, "agent," refer to the orange throat and parallel orange lines on the venter.

tamaulipensis: The name is derived from the name of the Sierra de Tamaulipas, and -ensis, a possessive suffix. 
References
  • Andrews, R.M., F.R. Méndez de la Cruz & M.V. Santa Cruz 1997. Body Temperature of Female Sceloporus grammicus: Thermal Stress or Impaired Mobility? Copeia 1997 (1): 108-115. - get paper here
  • ARÉCHAGA-OCAMPO, Samuel; Carlos A. MONTALBÁN HUIDOBRO & Rubén CASTRO-FRANCO 2008. NUEVOS REGISTROS Y AMPLIACION DE LA DISTRIBUCION DE ANFIBIOS Y REPTILES EN EL ESTADO DE MORELOS, MÉXICO. Acta Zoológica Mexicana (n.s.) 24 (2): 231-233 - get paper here
  • Arevalo, E.; Davis, S.K.;Sites, J.W. 1994. Mitochondrial DNA sequence divergence and phylogenetic relationships among eight chromosome races of the Sceloporus grammicus complex (Phrynosomatidae) in central Mexico. Systematic Biology 43 (3): 387-418 - get paper here
  • Arevalo, Elisabeth; Casas, Gustavo; Davis, Scott K.; Lara, Guillermo; Sites, Jack W., Jr 1993. Parapatric hybridization between chromosome races of the Sceloporus grammicus complex (Phrynosomatidae): Structure of the Ajusco transect. Copeia 1993 (2): 352-372 - get paper here
  • Baird, S.F. and Girard,C. 1852. Characteristics of some new reptiles in the Museum of the Smithsonian Institution, part 2. Proc. Acad. Nat. Sci. Philadelphia 6: 125-129 - get paper here
  • Bartlett, R. D. & Bartlett, P. 1999. A Field Guide to Texas Reptiles and Amphibians. Gulf Publishing Co., Houston, Texas, 331 pp.
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174 - get paper here
  • Blatchley, W.S. 1893. On a collection of batrachians and reptiles from Mount Orizaba, Mexico, with descriptions of two new species. Proc. US Natl. Mus. 16: 37-42 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger, G.A. 1897. A revision of the lizards of the genus Sceloporus. Proc. Zool. Soc. London 1897: 474-522 - get paper here
  • CANALES-GORDILLO, BRASIL; HIBRAIM ADÁN PÉREZ-MENDOZA, MIGUEL ADRIÁN DOMÍNGUEZ-GODOY, VÍCTOR ARGAEZ, ISRAEL SOLANO-ZAVALETA, ANÍBAL H. DÍAZ DE LA VEGA-PÉREZ & J. JAIME ZÚÑIGA-VEGA. 2022. Morphotypic Variation in Throat Coloration: A Continuous or a Discrete Trait? A Survival Analysis on a Polymorphic Lizard. Ichthyology & Herpetology 110(2): 388–397. - get paper here
  • Canseco-Márquez, L., & Gutiérrrez-Mayén, M.G. 2010. Anfibios y reptiles del Valle de Tehuacán-Cuicatlán. Comisión Nacional para el conocimiento y uso de la biodiversidad, México D.F., Mexico, 302 pp - get paper here
  • Carbajal-Márquez, Rubén A. and Gustavo E. Quintero-Díaz 2016. The Herpetofauna of Aguascalientes, México. Revista Mexicana de Herpetología 2(1):
  • Castro-Franco, Rubén, María Guasalupe Bustos-Zagal 1994. List of reptiles of Morelos, Mexico, and their distribution in relation to vegetation types. Southwestern Naturalist 39 (2): 171-175 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Cope, E.D. 1867. Fifth contribution lo the herpetology of tropical America. Proc. Acad. nat. Sci. Philadelphia, 18 [1866]: 317-323 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Cruz-Elizalde R, Ramírez-Bautista A, Pineda-López R, Mata-Silva V, DeSantis DL, García-Padilla E, Johnson JD, Rocha A, Fucsko LA, Wilson LD. 2022. The herpetofauna of Querétaro, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 16(1) [General Section]: 148–192 (e308) - get paper here
  • Cuellar, O., F.R. Méndez De La Cruz, M. Villagrán-Santa Cruz & R. Sánchez-Trejo 1996. Pregnancy does not increase the risk of mortality in wild viviparous lizards (Sceloporus grammicus) Amphibia-Reptilia, 17(1): 77-80. - get paper here
  • Degnan, P.H. and E. Arévalo 2004. Isolation of Microsatellite Loci in Sceloporus grammicus (Squamata, Phrynosomatidae). Americ. J. Undergrad. Res. 2 (4): 1-11 - get paper here
  • DÍAZ-MARÍN, CÉSAR; ABRAHAM LOZANO & AURELIO RAMÍREZ-BAUTISTA. 2022. Life history variation in four populations of the Graphic Spiny Lizard, Sceloporus grammicus (Squamata: Phrynosomatidae), in central Mexico. Salamandra 58(4): 289–301. - get paper here
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Dixon, James R. and Julio A. Lemos-Espinal 2010. Amphibians and reptiles of the state of Queretaro, Mexico. Tlalnepantla UNAM, 428 pp.
  • DOMINGUEZ GODOY, M. A., J. E. GOMEZ-CAMPOS, AND A. H. D. DE LA VEGA-PEREZ 2020. Sceloporus grammicus (Mesquite Lizard). Unusual Grouping And Mortality. Herpetological Review 51: 603-604.
  • DOMÍNGUEZ-GODOY, M.A.; R. HUDSON, B. MONTOYA, E. BASTIAANS & A.H. DÍAZ DE LA VEGA-PÉREZ. 2022. Too cool to fight: Is ambient temperature associated with male aggressive behavior in the mesquite lizard? Journal of Zoology 317(4): 283–293. - get paper here
  • DOMÍNGUEZ-GODOY, MIGUEL A.; ROBYN HUDSON, HIBRAHIM A. PÉREZ-MENDOZA, SERGIO ANCONA & ANÍBAL H. DÍAZ DE LA VEGA-PÉREZ. 2020. Living on the edge: Lower thermal quality but greater survival probability at a high altitude mountain for the mesquite lizard (Sceloporus grammicus). Journal of Thermal Biology 94: 102757. - get paper here
  • Dosselman, Dennis J.;Schaalje, G. Bruce;Sites, Jack W., Jr 1998. An analysis of fluctuating asymmetry in a hybrid zone between two chromosome races of the Sceloporus grammicus complex (Squamata: Phrynosomatidae) in central Mexico. Herpetologica 54 (4): 434-447 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Dunn, Emmett Reid 1936. The amphibians and reptiles of the Mexican Expedition of 1934. Proceedings of the Academy of Natural Sciences of Philadelphia 88: 471-477 - get paper here
  • Fernández-Badillo, Leonardo & Irene Goyenechea-Mayer Goyenechea 2010. Anfibios y reptiles del valle del Mezquital, Hidalgo, México. Revista Mexicana de Biodiversidad 81: 705- 712
  • Fernández-Badillo, Leonardo; Norma Leticia Manríquez-Morán; Jesús Martín Castillo-Cerón & Irene Goyenechea 2016. Análisis herpetofaunístico de la zona árida del estado de Hidalgo. Revista Mexicana de Biodiversidad 87: 156–170 - get paper here
  • Fitch, H. S. 1978. Sexual size differences in the genus Sceloporus. Univ. Kansas Sci. Bull. 51 (13): 441-461 - get paper here
  • GARCÍA-PADILLA, ELÍ; IVÁN VILLALOBOS- JUÁREZ & DAVID LAZCANO 2024. APUNTES SOBRE LA PRESENCIA DE LA VÍBORA DE CASCABEL DE LAS ROCAS TAMAULIPECA (CROTALUS MORULUS) EN TAMAULIPAS Biología y Sociedad 7 (13): 22-37
  • Gloyd, Howard K.;Smith, Hobart M. 1942. Amphibians and reptiles from the Carmen Mountains, Coahuila. Bulletin of the Chicago Academy of Sciences 6 (13): 231-235
  • Gonzalez Alonso, Arturo 1991. Distribucion geografica de una raza cromosomica del complejo Sceloporus grammicus (Sauria: Phrynosomatidae) en el Este de Mexico Central. Bol. Soc. Herpetol. Mex. 3 (1): 7-10 - get paper here
  • Goyenechea, Irene;Mendoza-Quijano, Fernando; Flores-Villela, Oscar; Reed, Kent M. 1996. Extreme chromosomal polytypy in a population of Sceloporus grammicus (Sauria: Phrynosomatidae) at Santuario Mapethé, Hildalgo, México. Journal of Herpetology 30 (1): 39-46 - get paper here
  • Gravenhorst, J. L. C. 1838. Beiträge zur genaueren Kenntniss einiger Eidechsengattungen. Nova Acta Acad. Caes. Leop.-Carol. 18: 712-784 [1837]
  • Gray, J. E. 1831. A synopsis of the species of Class Reptilia. In: Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed [V Whittaker, Treacher and Co., London: 481 + 110 pp. [1830] - get paper here
  • Guillette, L.J., Jr. & G. Casas-Andreu 1980. Fall Reproductive Activity in the High Altitude Mexican Lizard, Sceloporus grammicus microlepidotus Journal of Herpetology 14 (2): 143-147. - get paper here
  • Guillette, L.J., Jr. & G. Casas-Andreu 1981. Seasonal Variation in Fat Body Weights of the Mexican High Elevation Lizard Sceloporus grammicus microlepidotus Journal of Herpetology 15 (3): 366-371. - get paper here
  • Hall, W.P. 2009. Chromosome Variation, Genomics, Speciation and Evolution in Sceloporus Lizards. Cytogenet Genome Res 127: 143-165 - get paper here
  • Hall, William P. and Robert K. Selander 1973. Hybridization of Karyotypically Differentiated Populations in the Sceloporus grammicus Complex (Iguanidae). Evolution 27 (2): 226-242 - get paper here
  • Heimes, P. 2022. LIZARDS OF MEXICO - Part 1 Iguanian lizards. Edition Chimaira, Frankfurt Am Main, 448 pp. - get paper here
  • Hernández-Salinas, Uriel, Aurelio Ramírez-Bautista, Adrian Leyte-Manrique and Geoffery R. Smith. 2010. Reproduction and sexual dimorphism in two populations of Sceloporus grammicus (Sauria: Phrynosomatidae) from Hidalgo, Mexico. Herpetologica 66 (1): 12-22 - get paper here
  • Illescas-Aparicio, M., Clark-Tapia, R., González-Hernández, A., Vásquez-Díaz, Pedro R. & Aguirre-Hidalgo, V. 2016. Diversidad y riqueza herpetofaunística asociada al bosque de manejo forestal y áreas de cultivo, en Ixtlán de Juárez, Oaxaca. Acta Zoológica Mexicana (n.s.) 32(3): 359-369 - get paper here
  • Köhler, G. & Heimes, P. 2002. Stachelleguane. Herpeton-Verlag (Offenbach), 174 pp.
  • Lara-Góngora, G. 1986. New distributional records for some Mexican reptiles and amphibians. Bull. Maryland Herp. Soc. 22 (2): 62-67 - get paper here
  • Lara-Góngoro, Guillermo 2004. A new species of Sceloporus (Reptilia, Sauria: Phrynosomatidae) of the grammicus complex from Chihuahua and Sonora, Mexico. Bull. Maryland Herp. Soc. 40(1): 1-41 - get paper here
  • Lara-Resendiz, Rafael A. and Aníbal H. De La Vega-Pérez. 2013. Sceloporus grammicus (mesquite lizard) selected body temperature. Herpetological Review 44 (2): 328-329 - get paper here
  • Lazcano D, Nevárez-de los Reyes M, García-Padilla E, Johnson JD, Mata-Silva V, DeSantis DL, Wilson LD. 2019. The herpetofauna of Coahuila, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 13(2) [General Section]: 31–94 (e189) - get paper here
  • Leaché, A.D. 2010. Species trees for spiny lizards (Genus Sceloporus): Identifying points of concordance and conflict between nuclear and mitochondrial data. Molecular Phylogenetics and Evolution 54: 162–171 - get paper here
  • Leaché, A.D. & J.W.Sites, Jr. 2009. Chromosome Evolution and Diversification in North American Spiny Lizards (Genus Sceloporus). Cytogenet Genome Res 127: 166-181 - get paper here
  • Lemos-Espinal JA, Smith GR 2020. A conservation checklist of the amphibians and reptiles of the State of Mexico, Mexico with comparisons with adjoining states. ZooKeys 953: 137-159 - get paper here
  • Lemos-Espinal JA, Smith GR 2020. A conservation checklist of the amphibians and reptiles of Mexico City, with comparisons with adjoining states. ZooKeys 951: 109-131 - get paper here
  • Lemos-Espinal JA, Smith GR 2020. A conservation checklist of the herpetofauna of Morelos, with comparisons with adjoining states. ZooKeys 941: 121-144 - get paper here
  • Lemos-Espinal JA, Smith GR, Gadsden-Esparza H, Valdez-Lares R, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of Durango, Mexico, with comparisons with adjoining states. ZooKeys 748: 65-87 - get paper here
  • Lemos-Espinal JA, Smith GR, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of San Luis Potosí, Mexico, with comparisons with adjoining states. ZooKeys 753: 83-106 - get paper here
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Coahuila, Mexico. Universidad Nacional Autonoma de Mexico, 550 pp.
  • Lemos-Espinal, Julio A. and James R. Dixon 2013. Amphibians and Reptiles of San Luis Potosí. Eagle Mountain Publishing, xii + 300 pp.
  • Lemos-Espinal, Julio A., Geoffrey R. Smith 2015. Amphibians and reptiles of the state of Hidalgo, Mexico. Check List 11 (3): 1642 - get paper here
  • Leyte-Manrique A, Mata-Silva V, Báez-Montes O, Fucsko LA, DeSantis DL, García-Padilla E, Rocha A, Johnson JD, Porras LW, Wilson LD. 2022. The herpetofauna of Guanajuato, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 16(2) [General Section: 133–180 (e321) - get paper here
  • Leyte-Manrique, Adrian; Abel Antonio Buelna-Chontal,<br>Miguel Alejandro Torres-Dıaz, Christian Berriozabal-Islas, and Carlos Alberto Maciel-Mata 2019. A Comparison of Amphibian and Reptile Diversity Between Disturbed and Undisturbed Environments of Salvatierra, Guanajuato, Mexico. Tropical Conservation Science 12: 1–12 - get paper here
  • Leyte-Manrique, Adrian; Efrén M. Hernández Navarro y Luis A. Escobedo-Morales 2015. Herpetofauna de Guanajuato: Un análisis histórico y contemporáneo de su conocimiento. Revista Mexicana de Herpetología 1(1): 1–14
  • Loc-Barragán JA, Smith GR, Woolrich-Piña GA, Lemos-Espinal JA 2024. An updated checklist of the amphibians and reptiles of Nayarit, Mexico with conservation status and comparison with adjoining States. Herpetozoa 37: 25-42 - get paper here
  • Marshall, J. C.; Sites, J. W. 2001. A comparison of nuclear and mitochondrial cline shapes in a hybrid zone in the Sceloporus grammicus complex (Squamata; Phrynosomatidae). Molecular Ecology 10 (2): 435-450 - get paper here
  • Marshall, Jonathon C.; Arevalo, Elisabeth; Benavides, Edgar; Sites, Joanne L.; Sites, Jack W., Jr. 2006. Delimiting species: comparing methods for Mendelian characters using lizards of the Sceloporus grammicus (Squamata: Phrynosomatidae) complex. Evolution 60 (5):1050-1065 - get paper here
  • Martin, Plul S. 1958. A biogeography of reptiles and amphibians in the Gomez Farias Region, Tamaulipas, Mexico. Miscellaneous publications, Museum of Zoology, University of Michigan (101): 1-102 + 7 plates - get paper here
  • Martínez‐Torres M, Sánchez‐Rivera UÁ, Cruz‐Cano NB, Castro‐Camacho YJ, Luis J, Medrano A. 2019. A non‐invasive method for semen collection and evaluation in small and medium size lizards. Reprod Dom Anim. 2019;54(Suppl. 4): 54–58 - get paper here
  • Martín-Regalado, N.; M. C. Lavariega, R. M. Gómez–Ugalde & C. Rodríguez–Pérez 2016. Anfibios y reptiles de la sierra de Cuatro Venados, Oaxaca, México. Arxius de Miscel·lània Zoològica, 14 (2016): 217–232 - get paper here
  • Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here
  • McCranie, J.R. & Wilson, L.D. 2001. The herpetofauna of the Mexican State of Aguascalientes. Courier Forschungsinstitut Senckenberg 230: 1-57
  • MONTAÑO-RUVALCABA, CARLOS; JACOBO REYES-VELASCO & JASON M. JONES 2022. Geographic distribution: SCELOPORUS GRAMMICUS (Graphic Spiny Lizard). MÉXICO: COLIMA Herpetological Review 53 (1): 78
  • Nevárez-de-los-Reyes, Manuel, David Lazcano, Elí García-Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The Herpetofauna of Nuevo León, Mexico: Composition, Distribution, and Conservation. Mesoamerican Herpetology 3 (3): 558–638 - get paper here
  • Ochoa-Ochoa, L. M. & O. Flores-Villela. 2006. Areas de diversidad yendemismo de la herpetofauna mexicana. UNAM-CONABIO, Mexico, D.F.: 211 pp - get paper here
  • Ortega, A. 1986. Fat Body Cycles in a Montane Population of Sceloporus grammicus Journal of Herpetology 20 (1): 104-108. - get paper here
  • Ortega, A. & R. Barbault 1984. Reproductive Cycles in the Mesquite Lizard Sceloporus grammicus Journal of Herpetology 18 (2): 168-175. - get paper here
  • Ortega, Alfredo; Ricardo Rodriguez, Lucina Hernandez and Robert Barbault. 1984. Cycles Journaliers d’Activité chez deux Espèces Sympatriques de Sceloporus (Iguanidae), S. grammicus et S. scalaris. Amphibia-Reptilia 5 (3-4): 347 - 354 - get paper here
  • Ortega-Rubio, A., G. Halffter, R. Barbault, A. Castellanos & F. Salinas 1999. Growth of Sceloporus grammicus in La Michilía Biosphere Reserve, Mexico Journal of Herpetology 33 (1): 123-126. - get paper here
  • Palacios-Aguilar, Ricardo 2020. UNA LISTA COMENTADA DE LAS ESPECIES DE ANFIBIOS Y REPTILES CON LOCALIDAD TIPO EN GUERRERO, MÉXICO. Rev. Latinoamer. Herp. 3 (2): 43-60
  • Palacios-Aguilar, Ricardo & OSCAR FLORES-VILLELA 2018. An updated checklist of the herpetofauna from Guerrero, Mexico. Zootaxa 4422 (1): 1-24 - get paper here
  • Pérez-Mendoza, Hibraim A.; J. Jaime Zúñiga-Vega, Yazmín H. Zurita-Gutiérrez, Juan Fornoni, Israel Solano-Zavaleta, Ana L. Hernández-Rosas, and Alejandro Molina-Moctezuma 2013. Demographic Importance of the Life-Cycle Components in Sceloporus grammicus. Herpetologica Dec 2013, Vol. 69, No. 4: 411-435 [erratum in 70(1): 122] - get paper here
  • Peterson, A.T. et al. 2004. A preliminary biological survey of Cerro Piedra Larga, Oaxaca, Mexico: Birds, mammals, reptiles, amphibians, and plants. Anales del Instituto de Biología, Universidad Nacional Autónoma de México, Serie Zoología 75(2): 439-466
  • Porter, C.A. 1988. Triploidy in the Lizard, Sceloporus grammicus Journal of Herpetology 22 (1): 112-115. - get paper here
  • Pruett, Jake Allen, Alison G. Ossip-Klein, Cuauhcihuatl Vital-Garcia, J. Jaime Zúñiga-Vega and Diana K. Hews. 2015. Sceloporus grammicus (mesquite lizard) pursuit deterrent signaling. Herpetological Review 46 (3): 437-438 - get paper here
  • Reed, K.M.;Greenbaum, I.F.; Sites, J.W. 1995. Dynamics of a novel chromosomal polymorphism within a hybrid zone between two chromosome races of the Sceloporus grammicus complex (Sauria, Phrynosomatidae). Evolution 49 (1): 48-60 - get paper here
  • Reed, K.M.;Greenbaum, I.F.; Sites, J.W. 1995. Cytogenetic analysis of chromosomal intermediates from a hybrid zone between two chromosome races of the Sceloporus grammicus complex (Sauria, Phrynosomatidae). Evolution 49 (1): 37-47 - get paper here
  • Reuter Cortés, Adrián 1994. Analisis morfologico y cariotipico de tres poblaciones de Sceloporus grammicus (Sauria: Iguanidae) en un gradiente altitudinal de la Sierra del Ajusco. Bol. Soc. Herpetol. Mex. 6 (1): 1-5 - get paper here
  • Roth-Monzón AJ, Mendoza-Hernández AA, Flores-Villela O. 2018. Amphibian and reptile biodiversity in the semi-arid region of the municipality of Nopala de Villagrán, Hidalgo, Mexico. PeerJ 6: e4202 - get paper here
  • Schmidt, K.P., and D. Owens. 1944. Amphibians and reptiles of northern Coahuila, Mexico. Field Museum of Natural History, Zoological Series. Chicago. 29: 97-115. - get paper here
  • Schmidt, Karl P.;Shannon, Frederick A. 1947. Notes on amphibians and reptiles of Michoacan, Mexico. Zoological Series of Field Museum of Natural History 31 (9): 63-85
  • Sites, J. W., Jr.; Basten, C. J.;Asmussen, M. A. 1996. Cytonuclear genetic structure of a hybrid zone in lizards of the Sceloporus grammicus complex (Sauria, Phrynosomatidae). Molecular Ecology 5 (3): 379-92 - get paper here
  • Sites, J.W. & Dixon, J.R. 1981. A new subspecies of the Iguanid lizard, Sceloporus grammicus, from Northeastern Mexico, with comments on its evolutionary implications and the status of S. g. disparilis. Journal of Herpetology 15 (1): 59-69 - get paper here
  • Sites, J.W. Jr. 1983. Chromosome evolution in the iguanid lizard Sceloporus grammicus. I. Chromosome polymorphisms. Evolution 37: 38-53 - get paper here
  • Sites, J.W., Jr. 1982. Morphological Variation within and among Three Chromosome Races of Sceloporus grammicus (Sauria: Iguanidae) in the North-Central Part of Its Range Copeia 1982 (4): 920-941. - get paper here
  • Sites, J.W., Jr., J.L. Camarillo, A. Gonzalez, F. Mendoza, L. Javier, M. Mancilla & G. Lara-Gongora 1988. Allozyme Variation and Genetic Divergence within and between three Cyrotypes of the Sceloporus grammicus complex (Sauria: Iguanidae) in Central Mexico Herpetologica 44 (3): 297-307 - get paper here
  • Sites, J.W., Jr., R.K. Chesser & R.J. Baker 1988. Population Genetic Structure and the Fixation of Chromosomal Rearrangements in Sceloporus grammicus (Sauria, Iguanidae): A Computer Simulation Study Copeia 1988 (4): 1043-1053. - get paper here
  • Sites, Jack W., Jr; Davis, Scott K.; Hutchinson, Delbert W.; Maurer, Brian A.; Lara, Guillermo 1993. Parapatric hybridization between chromosome races of the Sceloporus grammicus complex (Phrynosomatidae): Structure of the Tulancingo transect. Copeia 1993 (2): 373-398 - get paper here
  • Sites, Jack W., Jr;Barton, Nicholas H.;Reed, Kent M. 1995. The genetic structure of a hybrid zone between two chromosome races of the Sceloporus grammicus complex (Sauria, Phrynosomatidae) in Central Mexico. Evolution 49 (1): 9-36 - get paper here
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
  • Smith, Hobart M. 1946. Handbook of Lizards: Lizards of the United States and of Canada. Comstock, Ithaca, NY, xxii + 557 pp.
  • Smith, Hobart M. & Laufe, Leonard E. 1945. Mexican amphibians and reptiles in the Texas Cooperative Wildlife Collections. Trans. Kansas Acad. Sci. 48: 325-354 - get paper here
  • Stejneger, L. 1916. A new lizard of the genus Sceloporus from Texas. Proc. Biol. Soc. Washington 29: 227-230 - get paper here
  • Stuart‐Fox, D., Aulsebrook, A., Rankin, K.J., Dong, C.M. and McLean, C.A. 2020. Convergence and divergence in lizard colour polymorphisms. Biological Reviews - get paper here
  • Tanner, W.W. 1987. Lizards and turtles of Western Chihuahua. Great Basin Naturalist. 47: 383-421 - get paper here
  • Taylor, Edward H. 1952. Third contribution of the herpetology of the Mexican state of San Luis Potosí. Univ. Kansas Sci. Bull. 34 (13): 793-815 - get paper here
  • Tepos-Ramírez M, Garduño-Fonseca FS, Peralta-Robles CA, García-Rubio OR, Cervantes Jiménez R 2023. Annotated checklist of amphibians and reptiles from Querétaro, Mexico, including new records, and comments on controversial species. Check List 19(2): 269-292 - get paper here
  • Terán-Juárez, Sergio A., Elí García Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The herpetofauna of Tamaulipas, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (1): 43–113 - get paper here
  • Torres-Hernández, LA, Ramírez-Bautista A, Cruz-Elizalde R, Hernández-Salinas U, Berriozabal-Islas C, DeSantis DL, Johnson JD, Rocha A, García-Padilla E, Mata-Silva V, Fucsko LA, and Wilson LD. 2021. The herpetofauna of Veracruz, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 15(2) [General Section]: 72–155 - get paper here
  • Valdez-Lares, R.; R. Muñiz-Martínez; E.Gadsden; G. Aguirre-León; G. Castañeda-Gaytán; R. Gonzalez-Trápaga 2013. Checklist of amphibians and reptiles of the state of Durango, México. Check List 9 (4):714-724 - get paper here
  • Van Devender, Thomas R.; Lowe, Charles H. Lowe, Jr. 1977. Amphibians and reptiles of Yepomera, Chihuahua, Mexico. Journal of Herpetology 11 (1): 41-50 - get paper here
  • Vázquez Díaz, Joel;Quintero Díaz, Gustavo E. 2005. Anfibios y Reptiles de Aguascalientes [2nd ed.]. CONABIO, CIEMA, 318 pp.
  • Vite-Silva, Victor D.; Aurelio Ramírez-Bautista y Uriel Hernández-Salinas 2010. Diversidad de anfibios y reptiles de la Reserva de la Biosfera Barranca de Metztitlán, Hidalgo, México Diversity of amphibians and reptiles from the Barranca de Metztitlán Biosphera Reserve in Hidalgo, Mexico. Revista Mexicana de Biodiversidad 81: 473- 485 - get paper here
  • Wiegmann,A.F.A. 1828. Beyträge zur Amphibienkunde. Isis von Oken 21 (4): 364-383 - get paper here
  • Wiegmann,A.F.A. 1834. Herpetologia Mexicana, seu descriptio amphibiorum novae hispaniae , quae itineribus comitis de Sack, Ferdinandi Deppe et Chr. Guil. Schiede im Museum Zoologicum Berolinense Pervenerunt. Pars prima, saurorum species. Berlin, Lüderitz, iv + 54 pp. - get paper here
  • Wiens, John J.; Caitlin A. Kuczynski, Saad Arif & Tod W. Reeder 2010. PHYLOGENETIC RELATIONSHIPS OF PHRYNOSOMATID LIZARDS BASED ON NUCLEAR AND MITOCHONDRIAL DATA, AND A REVISED PHYLOGENY FOR SCELOPORUS. Molecular Phylogenetics and Evolution 54: 150-161 - get paper here
  • Wilson, Larry David;McCranie, James R. 1979. Notes on the herpetofauna of two mountain ranges in Mexico (Sierra Fria, Aguascalientes, and Sierra Morones, Zacatecas. Journal of Herpetology 13 (3): 271-278 - get paper here
  • Woolrich-Piña, G. A., E. García-Padilla, D. L. DeSantis, J. D. Johnson, V. Mata-Silva, and L. D. Wilson 2017. The herpetofauna of Puebla, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 4(4): 791–884 - get paper here
  • Woolrich-Piña, Guillermo A, Paulino Ponce-Campos, Jesús Loc-Barragán, Juan Pablo Ramírez-Silva, Vicente Mata-Silva, Jerry D. Johnson, Elí García-Padilla and Larry David Wilson. 2016. The herpetofauna of Nayarit, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (2): 376-448 - get paper here
  • Zúñiga-Vega, J Jaime, Jake A Pruett, Alison G Ossip-Drahos, Stephanie M Campos, Ryan J Seddon, Savannah L Price, Cristina Romero-Diaz, et al. 2021. Information out of the Blue: Phenotypic Correlates of Abdominal Color Patches in Sceloporus Lizards. Zoology 149: 125961 - get paper here
  • Zúñiga-Vega, J. J., C. Molina-Zuluaga, et al. 2012. Relative importance of reproductive life-history paths in one population of the lizard Sceloporus grammicus. Amphibia-Reptilia 33 (3-4): 401-413 - get paper here
  • Zúñiga-Vega, Jaime J.; Méndez-de la Cruz, Fausto R.; Cuellar, Orlando 2008. Body growth in one montane population of Sceloporus grammicus (Sauria, Phrynosomatidae) in Central México. Amphibia-Reptilia 29 (1): 127-134 - get paper here
 
External links  
Is it interesting? Share with others:

As link to this species use URL address:

https://reptile-database.reptarium.cz/species?genus=Sceloporus&species=grammicus

without field 'search_param'. Field 'search_param' is used for browsing search result.



Please submit feedback about this entry to the curator