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Smaug vandami (FITZSIMONS, 1930)

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Higher TaxaCordylidae (Cordylinae), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Van Dam’s Girdled Lizard 
SynonymZonurus vandami FITZSIMONS 1930: 25
Zonurus vandami perkoensis FITZSIMONS 1930: 27
Cordylus vandami vandami — FITZSIMONS 1943
Cordylus warreni perkoensis — LOVERIDGE 1944
Cordylus warreni vandami — LOVERIDGE 1944
Cordylus vandami — BRANCH 1998
Smaug vandami — STANLEY et al. 2011
Smaug vandami — REISSIG 2014 
DistributionRepubic of South Africa (North Province)

Type locality: Gravelotte, near Leydsdorp, Eastern Transvaal

perkoensis: Type locality: on farm “Perkoe”, near the Olifants River, Eastern Transvaal.  
ReproductionOvoviviparous. 
TypesHolotype: DNMNH (= TM) 7407
Holotype: DNMNH (= TM) 12804, adult female [perkoensis] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2648 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSee also S. warreni.

Synonymy partly after JACOBSEN (unpubl. thesis, 1989) and E. Stanley (pers. comm., 31 Oct 2011). 
Etymologynamed after the collector of the types, G.P.F. van Dam, Esq. 
References
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Branch, W. R. 1998. Field Guide to the Snakes and Other Reptiles of Southern Africa. 3rd ed. Fully Revised and Updated to Include 83 New Species. Ralph Curtis Books (Sanibel Island, Florida), 399 pp.
  • FitzSimons, V. F. 1930. Descriptions of new South African Reptilia and Batrachia, with distribution records of allied species in the Transvaal Museum collection. Annals Transvaal Mus. 14: 20-48. - get paper here
  • FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp.
  • Harport, V. 2019. Familienurlaub mal anders – Südafrikas Norden. Reptilia (Münster) 24 (135): 70-79 - get paper here
  • Loveridge,A. 1944. Revision of the African lizards of the family Cordylidae. Bull. Mus. comp. Zool. Harvard 95 (1): 1-118 - get paper here
  • Mouton, le Fras; Alexander Flemming; Michael Bates and Chris Broeckhoven 2018. The relationship between generation gland morphology and armour in Dragon Lizards (Smaug): a reassessment of ancestral states for the Cordylidae. Amphibia-Reptilia 39 (4): 445–456 - get paper here
  • Reissig, J. 2014. Girdled Lizards and Their Relatives. Edition Chimaira, Frankfurt, ISBN-10: 3899734378
  • Stanley, E. L. and Bates, M. F. 2014. Here be dragons: a phylogenetic and biogeographical study of the Smaug warreni species complex (Squamata: Cordylidae) in southern Africa. Zoological Journal of the Linnean Society 172 (4): 892–909; doi: 10.1111/zoj.12187 - get paper here
  • Stanley, Edward L.; Aaron M. Bauer; Todd R. Jackman, William R. Branch, P. Le Fras N. Mouton 2011. Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): 53-70. - get paper here
 
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