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Stenocercus formosus (TSCHUDI, 1845)

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Higher TaxaTropiduridae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Formosa Whorltail Iguana 
SynonymScelotrema formosum TSCHUDI 1845: 155
Liocephalus formosus BOULENGER 1880: 43
Liocephalus formosus — BOULENGER 1885: 168
Liocephalus rhodogaster BOULENGER 1901: 547
Liocephalus lineogularis WERNER 1901: 3
Stenocercus seydi ANDERSSON 1908
Leiocephalus lineogularis — BURT & BURT 1931
Leiocephalus seydi — BURT & BURT 1933
Leiocephalus formosus — BURT & BURT 1933
Ophryoessoides formosus — ETHERIDGE 1966: 88
Ophryoessoides lineogularis — ETHERIDGE 1966: 88
Ophryoessoides arenarius — PETERS et al. 1970: 213
Ophryoessoides formosus — PETERS et al. 1970: 214
Stenocercus formosus — FRITTS 1974
Stenocercus formosus — CEI 1993
Stenocercus formosus — TORRES-CARVAJAL 2004
Stenocercus formosus — TORRES-CARVAJAL 2007 
DistributionPeru (Junin)

Type locality: La Merced, 1000 m elevation, Peru; Lectotype from “mountains of central Peru, Río Tulumayo, Junín, Peru”.

rhodogaster: Type locality: “Merced, Perené River, 3250 feet”.

lineogularis: Type locality: “Chanchamayo”.

seydi: Type locality: “Peru, La Merced”.  
Reproductionoviparous 
TypesLectotype: MHNN 2266 (Neuchatel, fide ORTIZ 1989)
Holotype: MWNH 473 [Stenocercus seydi]
Syntypes: BMNH 1900.11.27.24–25 (= 1946.8.29.81–82) [rhodogaster]
Holotype: lost, was MTD 1781 [lineogularis] 
DiagnosisDiagnosis in TORRES-CARVAJAL 2007. 
CommentSynonymy: fide Carvalho 2021.

Distribution: see partial map (Peru) in Venegas et al. 2020: 563. Does not occur in Ecuador as stated by previous authors (e.g. PETERS & DONOSO-BARROS 1970).

Belongs to the clade Anatomegalepis (fide TORRES-CARVAJAL 2006). 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger, George A. 1880. Reptiles et Batraciens recueillis par M. Emile de Ville dans les Andes de l'équateur. Bulletin de la Société Zoologique de France 5: 41-48 - get paper here
  • Carranza, S.; E. N. Arnold & F. Amat 2004. DNA phylogeny of Lacerta (Iberolacerta) and other lacertine lizards (Reptilia: Lacertidae): did competition cause long-term mountain restriction? Systematics and Biodiversity 2 (1): 57-77 - get paper here
  • Carvalho, André L.G. 2021. Resolving the Obscure Identity of Steironotus arenarius Tschudi, 1845 and Tropidurus tschudii Roux, 1907 (Squamata: Tropiduridae) South American Journal of Herpetology 19(1), 40-53 - get paper here
  • Etheridge, Richard E. 1966. The systematic relationships of West Indian and South American lizards referred to the iguanid genus Leiocephalus. Copeia 1966 (1): 79-91 - get paper here
  • Fritts, Thomas H. 1974. A multivariate evolutionary analysis of the Andean Iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1-89
  • Köhler, J. & Güsten, R. 2007. Herpetological type specimens in the natural history collections of the museums in Darmstadt and Wiesbaden, Germany. Spixiana 30 (2): 275-288 - get paper here
  • Ortiz J C 1989. Catalogue des types du musée d’histoire naturelle de Neuchatel III. Sauriens. Bulletin de la Societe neuchateloise des Sciences naturelles 112: 47-64 - get paper here
  • Torres-Carvajal, O. 2004. The abdominal skeleton of Tropidurid lizards (Squamata: Tropiduridae). Herpetologica 60 (1): 75-83 - get paper here
  • Torres-Carvajal, Omar; James A. Schulte II and John E. Cadle 2006. Phylogenetic relationships of South American lizards of the genus Stenocercus (Squamata: Iguania): A new approach using a general mixture model for gene sequence data. Molecular Phylogenetics and Evolution 39 (1): 171-185 - get paper here
  • VENEGAS, PABLO J.; LOURDES Y. ECHEVARRÍA, LUIS A. GARCÍA-AYACHI, CAROLL Z. LANDAURO 2020. Two new sympatric species of Stenocercus (Squamata: Iguania) from the inter-Andean valley of the Mantaro River, Peru. Zootaxa 4858 (4): 555–575 - get paper here
 
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