You are here » home › search results › Testudo hermanni

Testudo hermanni GMELIN, 1789

IUCN Red List - Testudo hermanni - Near Threatened, NT
iNaturalist logo

Can you confirm these amateur observations of Testudo hermanni?

Add your own observation of
Testudo hermanni »

Find more photos by Google images search: Google images

Higher TaxaTestudinidae, Testudinoidea, Testudines (turtles)
SubspeciesTestudo hermanni hermanni GMELIN 1789
Testudo hermanni boettgeri MOJSISOVICS 1889 
Common NamesE: Hermann’s Tortoise
G: Griechische Landschildkröte
E: Eastern Hermann’s Tortoise [boettgeri]
E: Western Hermann’s Tortoise [hermanni]
Croatian: kopnena kornjača
Italian: testuggine di Hermann 
SynonymTestudo hermanni GMELIN in LINNAEUS 1789
Testudo graeca bettai LATASTE 1881
Testudo graeca — BOULENGER 1889
Testudo hermanni robertmertensi WERMUTH 1952
Testudo hermanni robertmertensi — WERMUTH & MERTENS 1977
Protestudo hermanni — CHKHIKVADZE 1970
Testudo hermanni robertmertensi — FORMAN — & FORMAN 1981
Testudo hermanni — ENGELMANN et al. 1993
Eurotestudo hermanni — LAPPARENT DE BROIN et al. 2006
Testudo (Chersine) hermanni — TTWG 2017: 157
Chersine hermanni — VLACHOS & RABI 2019

Testudo hermanni hermanni GMELIN 1789
Testudo graeca var. hercegovinensis WERNER 1899
Testudo enriquesi PARENZAN 1932
Testudo hermanni hermanni — WERMUTH 1952
Testudo hercegovinensis — PERÄLÄ 2001
Testudo hermanni hermanni — KUYL et al. 2002
Testudo hercegovinensis — VINKE & VINKE 2004
Testudo hermanni hercegovinensis — ROGNER 2005
Testudo hercegovinensis — BONIN et al 2006
Testudo hermanni hercegovinensis — PAWLOWSKI & KRÄMER 2009
Testudo hermanni hercegovinensis — WIRTH 2010
Testudo hermanni hercegovinensis — WIRTH 2012
Testudo (Chersine) hermanni hermanni — TTWG 2017: 157
Testudo (Chersine) hermanni hermanni — TTWG 2021

Testudo hermanni boettgeri MOJSISOVICS 1889
Testudo graeca var. boettgeri MOJSISOVICS 1889: 242 [non Testudo boettgeri SIEBENROCK 1904]
Testudo hermanni boettgeri — ENGELMANN et al. 1993
Testudo hermanni boettgeri — KUYL et al. 2002
Testudo hermanni boetgeri — TÜRKOZAN et al. 2005 (in error)
Eurotestudo boettgeri — LAPPARENT DE BROIN et al. 2006
Testudo boettgeri — BONIN et al 2006
Eurotestudo boettgeri — ROULIN et al. 2013
Testudo hermanni boettgeri — KOPPITZ et al. 2017
Testudo (Chersine) hermanni boettgeri — TTWG 2017: 157
Testudo (Chersine) hermanni boettgeri — TTWG 2021 
DistributionE Spain (also introduced to the Balearics: Mallorca, Menorca),
S France (incl. Corsica),
Italy (incl. Sardinia, Sicily, Elba, Pianosa, Lampedusa island),
Albania, Croatia (including some Adriatic islands),
Bosnia and Hercegovina, Montenegro, Macedonia, Serbia,
Bulgaria, Romania,
Greece (incl. the Ionian Islands, Corfu = Corfou)
E Turkey

boettgeri: coastal regions of the former Yugoslavia, through Albania, Greece, Bulgaria, to the Black Sea. Type locality: Orsova, Banat.

Type locality: unknown (fide WERMUTH & MERTENS 1977)  
Reproductionoviparous. Temperature‐dependent sex determination (TSD) fide Nemesházi & Bókony 2023. 
TypesHolotype: MZUS 121 (Musée Zoologique de la ville de Strasbourg)
Holotype: SMF 7836 [boettgeri]
Holotype: SMF 37468 [robertmertensi] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2386 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSubspecies: Testudo hermanni robertmertensi has a yellow patch behind the eye which T. h. hermanni doesn't have. However, Highfield (1992) and others consider robertmertensi as invalid. Eastern populations should be called T. h. boettgeri in Highfield’s opinion.

Phylogeny: PARHAM et al. (2005, 2006) showed that T. hermanni is nested in a clade with Indotestudo, Agrionemys and Malacochersus and thus paraphyletic with respect to other Testudo species. In order to resolve the paraphyly “Testudo” hermanni may be placed in the genus “Agrionemys”.

Synonymy: Eurotestudo Lapparent de Broin et al., 2006, which was recently erected with the type species T. hermanni, is an objective junior synonym of Chersine Merrem, 1820 and Medaestia Wussow, 1916 according to FRITZ & BININDA-EMONDS (2007).

Hybridization: T. horsfieldi and T. hermanni may hybridize (Kirsche 1984).

Distribution: For a map see Escoriza & Hassine 2022: Fig. 1. 
EtymologyNamed after Jean Hermann (1738-1800), Professor of Natural History and herpetologist in Strasbourg, France. 
References
  • Amiranashvili, N.G. 2000. Differences in shell morphology of Testudo graeca and Testudo hermanni , based on material from Bulgaria. Amphibia-Reptilia 21 (1): 67-81 - get paper here
  • Arsovski, D. , Olivier, A. , Bonnet, X. , Drilholle, S. , Tomović, L. , Béchet, A. , Golubović, A. and Besnard, A. 2018. Covariates streamline age‐specific early life survival estimates of two chelonian species. J Zool, 306: 223-234. doi:10.1111/jzo.12585 - get paper here
  • Auffenberg, Walter 1977. Display Behavior in Tortoises. American Zoologist 17: 241-250 - get paper here
  • Badiane, Arnaud; Cátia Matos and Xavier Santos 2017. Uncovering environmental, land-use and fire effects on the distribution of a low-dispersal species, the Hermann’s tortoise Testudo hermanni. Amphibia-Reptilia 38 (1): 67 - 77 - get paper here
  • Baur, Markus 2013. Zehn Fragen zur Ernährung von Landschildkröten. Draco 13 (54): 68-79 - get paper here
  • BECH, N.; D. NIVELLE, S. CARON, J.M. BALLOUARD, V. ARNAL, D. ARSOVSKI, A. GOLUBOVIĆ, X. BONNET & C. MONTGELARD. 2022. Extent of introgressive hybridization in the Hermann’s tortoise (Testudo hermanni hermanni) from the south of France. European Journal of Wildlife Research 68(3): 37. - get paper here
  • Bender, C. & Henle, K. 2007. Kennzeichnung von Landschildkröten der Gattung Testudo: Richtlinien für die Fotodokumentation von Jungtieren - Kurzfassung. Elaphe 15 (1): 51-54
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bertolero, A. & A. Marín 2005. Efficacy of inguinal palpation for detecting oviductal eggs in Hermann's tortoise, Testudo hermanni Amphibia-Reptilia, 26(4): 523-526. - get paper here
  • Bertolero, A., M.A. Carretero & G. A. Llorente 2005. An assessment of the reliability of growth rings counts for age determination in the Hermann's Tortoise Testudo hermanni Amphibia-Reptilia, 26(1): 17-23. - get paper here
  • Bertolero, Albert. 2006. La Tortue d'Hermann Testudo hermanni sur les îles de Majorque et de Minorque. Chéloniens 1: 12-18
  • Bertolero, Albert; Nougarède, Jean Pierre; Cheylan, Marc; Marín, Aurelio 2007. Breeding traits of Hermann's tortoise Testudo hermanni hermanni in two western populations. Amphibia-Reptilia 28 (1): 77-85 - get paper here
  • Bertolero, Albert; Nougarède, Jean-Pierre; Cheylan, Marc 2007. Female reproductive phenology in a population of Hermann's tortoise Testudo hermanni hermanni in Corsica. The Herpetological Journal 17: 92-96 - get paper here
  • Bertolero, Albert;MartØnez-Vilalta, Albert 1994. Presencia histórica de Testudo hermanni en las comarcas del Baix Ebre y Montsié (sur de Cataluña). Boletín de la Asociación Herpetológica Española 1994 (5): 2-3 - get paper here
  • Beshkov, V & Nanev, K 2006. The Amphibians and Reptiles in Bulgaria. Pensoft Series Faunistica 46, 120 pp.
  • Biaggini, Marta and Claudia Corti 2018. Facing Habitat Reduction in Your Own Shell: Patterns of Non-lethal Injuries in the Endangered Tortoise Testudo hermanni in Italy. Herp. Cons. Biol. 13 (3): 539–550 - get paper here
  • Biaggini, Marta; Antonio Romano, Luciano Di Tizio, Claudia Corti 2018. Density and sex-ratio of wild populations of three Testudo species in Italy. Herpetozoa 30 (3/4): 203 - 208 - get paper here
  • Bonin, F., Devaux, B. & Dupré, A. 2006. Turtles of the World. English translation by P.C.H. Pritchard. Johns Hopkins University Press, 416 pp.
  • Bour, R. 2004. A new character for the identificaiton of populations of the Hermann’s tortoise, Testudo hermanni GMELIN, 1789 (Chelonii, Testudinidae). Salamandra 40 (1): 59-66 - get paper here
  • Bour, Roger; Ohler, Annemarie. 2008. Chersine Merrem, 1820 and Chersina Gray, 1831: a nomenclatural survey. Zootaxa 1752: 66-68 - get paper here
  • Brugnola, Luca, Carlo Biancardi, Nicoletta Di Francesco, Luciano Di Tizio and Adrian Gheorghiu. 2013. DataDotDNA: an alternative marking system for tortoises of the genus Testudo. Acta Herpetologica 8 (2): 163-166 - get paper here
  • Buchert, Peter B 2012. Eine neue Heimat für Landschildkröten. Draco 12 (49): 4-17
  • Budó, Joan; Xavier Capalleras, Jenar Fèlix, & Joan Font 2009. Aportacions sobre l’estudi de l’alimentació de la tortuga mediterrània Testudo hermanni hermanni (Gmelin, 1789) a la serra de l’Albera (Catalunya). Butll. Soc. Cat. Herp. 18: 109-115
  • Causa, Maria Casamitjana; Juan Carlos Loaiza Usuga, Núria Simon Bartomeus, Pere Frigola Vidal 2012. Aspectos ecológicos y efectos del manejo forestal en una población de tortuga mediterránea (Testudo hermanni hermanni) en Cataluña (España). Basic and Applied Herpetology 26: 73-86 - get paper here
  • Chelazzi, G. & G. Delfino 1986. A Field Test on the Use of Olfaction in Homing by Testudo hermanni (Reptilia: Testudinidae) Journal of Herpetology 20 (3): 451-455. - get paper here
  • Chessman, B. C. 2024. The conundrum of turtle and tortoise basking: A critical review. Journal of Zoology - get paper here
  • Cheylan, M. 1984. The true Status and Future of Hermann’s Tortoise Testudo hermanni robertmertensi Wermuth 1952 in Western Europe Amphibia-Reptilia, 5(1): 17-26. - get paper here
  • Cimatti, E. 2006. Testudo hermanni - Hermann’s tortoise, questions and answers. Reptilia (GB) (47): 47-52 - get paper here
  • Cogălniceanu, Dan; Laurentiu Rozylowicz, Paul Székely, Ciprian Samoilă, Florina Stănescu, Marian Tudor, Diana Székely, Ruben Iosif 2013. Diversity and distribution of reptiles in Romania. ZooKeys 341 (2013): 49-76 - get paper here
  • Corti, C., Biaggini, M., Nulchis, V., Cogoni, R., Cossu, I. M., Frau, S., ... & Bassu, L. 2022. Species diversity and distribution of amphibians and reptiles in Sardinia, Italy. Acta Herpetologica, 17(2), 125-133
  • Corti, Claudia & Marco A.L. Zuffi 2003. Aspects of population ecology of Testudo hermanni hermanni from Asinara Island, NW Sardinia (Italy, Western Mediterranean Sea): preliminary data. Amphibia-Reptilia 24 (4): 441-447 - get paper here
  • Couturier, Thibaut; Cheylan, Marc; Guérette, Emmanuelle; Besnard, Aurélien 2011. Impacts of a wildfire on the mortality rate and small-scale movements of a Hermann's tortoise Testudo hermanni hermanni population in southeastern France. Amphibia-Reptilia 32 (4): 541-545 - get paper here
  • Cutuli, G.; M. Vannini, S. Fratini 2014. Demographic structure and genetic variability of a population of Testudo hermanni hermanni (Reptilia: Testudines: Testudinidae) from Southern Tuscany (Central Italy): a case of “happy-ending” uncontrolled reintroduction. Italian Journal of Zoology Vol. 80, Iss. 4, 2013 - get paper here
  • Daubner, M. & Vinke, T. 2006. Schildkröten im Fokus - Sonderband Testudo - häufig gehaltene Arten. Dauvi-Verlag, Bergheim, 157 pp.
  • Del Vecchio, S., R.L. Burke, L. Rugiero, M. Capula & L. Luiselli 2011. Seasonal Changes in the Diet of Testudo hermanni hermanni in Central Italy Herpetologica 67 (3): 236-249. - get paper here
  • Delaugerre, M., & Cheylan, M. 1992. Atlas de répartition des batraciens et reptiles de Corse. Ajaccio Parc naturel régional de Corse: Ecole pratique des hautes études, 128 pp. - get paper here
  • Djordjevíc, Sonja; Djurakíc, Marko; Golubovíc, Ana; Ajtíc, Rastko; Tomovíc, Ljiljana; Bonnet, Xavier 2011. Sexual body size and body shape dimorphism of Testudo hermanni in central and eastern Serbia. Amphibia-Reptilia 32 (4): 445-458 - get paper here
  • DOMINGO OLLÉ, ROGER; IKER A. SEVILLA, CARLES JUAN-SALLÉS, JOSEBA M. GARRIDO & JORDI JIMÉNEZ SANTAMARÍA. 2022. Mycobacterial Cloacitis with Systemic Dissemination in a Hermann’s Tortoise (Testudo hermanni). Journal of Herpetological Medicine and Surgery. - get paper here
  • Drechsler, Robby; Mercè Vilalta and Juan S. Monrós 2016. Analysis of movement patterns and macrohabitat use in Hermann’s tortoises (Testudo hermanni hermanni, Gmelin 1789) reintroduced in a coastal area dominated by pinewood in eastern Spain. Amphibia-Reptilia, DOI: 10.1163/15685381-00003065 - get paper here
  • Duro, S., Sönmez, B., Gündemir, O., Jashari, T., & Szara, T. 2021. Morphological Divergence of Hermann’s Tortoise (Testudo hermanni boettgeri Mojsisovits, 1889) in Albania. Animals 11 (1), 134 - get paper here
  • Eendebak, B. 2003. Een gezond, duurzame groep Griekse Landschildpadden (Testudo hermanni boettgeri). Lacerta 61 (3): 96-101 - get paper here
  • Ehrengart, W. 1971. Zur Pflege und Zucht der Griechischen Landschildkröte (Testudo h. hermanni). Salamandra 7 (2): 71-80 - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Escoriza D & J. Ben Hassine 2022. Niche diversification of Mediterranean and southwestern Asian tortoises. PeerJ 10:e13702 - get paper here
  • Esterbauer, H. 2019. Haltung, Zucht und Aufzucht der Östlichen Griechischen Landschildkröte, Testudo hermanni boettgeri, in einer Freianlage. Reptilia (Münster) 24 (136): 44-50 - get paper here
  • Forman, F. & Forman, B. 1981. Herpetologische Beobachtungen auf Korsika. Herpetofauna 3 (10): 12-16 - get paper here
  • Frank, Thomas; Enerit Saçdanaku, Michael Duda & Ferdinand Bego 2018. Amphibian and reptile fauna of the Vjosa River, Albania. Acta ZooBot Austria 155: 323–336 - get paper here
  • Fritz, Uwe and Olaf R.P. Bininda-Emonds 2007. When genes meet nomenclature: Tortoise phylogeny and the shifting generic concepts of Testudo and Geochelone. Zoology 110 (4): 298-307 [erratum in vol. 111 (1): 84] - get paper here
  • Fritz, Uwe; Auer, Markus; Bertolero, Albert; Cheylan, Marc; Fattizzo, Tiziano; Hundsdorfer, Anna K.; Martin Sampayo, Marcos; Pretus, Joan L.; Siroky, Pavel, Wink, Michael 2006. A rangewide Phylogeography of Hermann’s tortoise, Testudo hermanni (Reptilia: Testudines: Testudinidae): implications for taxonomy. Zoologica Scripta 35 (5):531-543. - get paper here
  • Galeotti, P., R. Sacchi, D.P. Rosa & M. Fasola 2007. Olfactory Discrimination of Species, Sex, and Sexual Maturity by the Hermann's Tortoise Testudo hermanni Copeia 2007 (4): 980-985. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Giacalone, Gabriele; Mario Lo Valvo; Uwe Fritz 2009. Phylogeographic link between Sicilian and Corso-Sardinian Testudo h. hermanni confirmed. Acta Herpetologica 4 (2): 119-123 - get paper here
  • Gmelin, J.F. 1789. Caroli a Linné Systema naturae. 13. ed., Tom 1 Pars 3. G. E. Beer, Lipsiae. 1033-1516 pp. - get paper here
  • Golubovic, Ana; Tomovic Ljiljana, Ivanovic Ana 2015. Geometry of self righting: the case of Hermann’s tortoises. Zoologischer Anzeiger 254: 99-105 - get paper here
  • Guyot, Ghislaine;Devaux, Bernard 1997. Variation in shell morphology and color of Hermann's Tortoise, Testudo hermanni, in southern Europe. Chelonian Conserv. Biol. 2 (3): 390-395
  • GVOZDENOVIĆ, S., & IKOVIĆ, V. 2022. CHECKLIST OF AMPHIBIANS AND REPTILES OF MONTENEGRO AND THEIR CONSERVATION STATUS. Natura Montenegrina, Podgorica, 2022, 14 (2): 93-105 - get paper here
  • Hailey, A., E.A. Pulford & D. Stubbs 1984. Summer Activity Patterns of Testudo hermanni Gmelin in Greece and France Amphibia-Reptilia, 5(1): 69-78. - get paper here
  • Hallmen, M. 2009. Bau einer Freianlage mit Gewächshaus zur ganzjährigen Haltung von Griechischen Landschildkröten (Testudo hermanni boettgeri) Terraria (16) 4 (2): 18-29.
  • Haxhiu, Idriz 1995. Current data on the chelonians of Albania. Chelonian Conserv. Biol. 1 (4): 326-327
  • Haxhiu, Idriz 1995. Results of studies on the chelonians of Albania. Chelonian Conserv. Biol. 1 (4): 324-326
  • Heimann, E. 1990. Testudo hermanni GMELIN, 1789. Sauria 12 Suppl.: 175–178 - get paper here
  • Heimann, E. 1992. Eine F2-Generation bei Testudo hermanni boettgeri MOJSISOVICS 1889 in Gefangenschaft. Sauria 14 (2): 19-22 - get paper here
  • Heimann, E. 2000. Eine F3-Generation bei Testudo hermanni boettgeri MOJSISOVICS, 1889 in menschlicher Obhut. Sauria 22 (3): 41-44 - get paper here
  • Henle, K. 1985. Ökologische, zoogeographische und systematische Bemerkungen zur Herpetofauna Jugoslawiens. Salamandra 21 (4): 229-251 - get paper here
  • Herz, M. 2002. Testudo hermanni boettgeri MOJSISOVICS 1889 in Norddalmatien. Sauria 24 (4): 19-22 - get paper here
  • Highfield A C 1989. Taxonomic diagnostic characters of tortoises (1): observations on the taxonomic significance of the division of the supracaudal scute in Testudo. British Herpetological Society Bulletin (30): 14-18
  • Highfield,A.C. 1992. Notes on the Systematics and Ecology of Testudo hermanni GMELIN 1789. Tortuga Gazette 28 (9): 1-3
  • Hitschfeld, Ellen; Auer, Markus; Fritz, Uwe 2008. Phalangeal formulae and ontogenetic variation of carpal morphology in Testudo horsfieldii and T. hermanni. Amphibia-Reptilia 29 (1): 93-99 - get paper here
  • Hout-Daubremont, C., C. Grenot & D. Bradshaw 1996. Temperature regulation in the tortoise Testudo hermanni, studied with indwelling probes Amphibia-Reptilia, 17(2): 91-102. - get paper here
  • Iftime, Alexandru 2002. Testudo hermanni Gmelin, 1789 in Dobroudja (SE Romania), with comments on conservation. Herpetozoa 15 (3/4):183-186. - get paper here
  • Iosif, Ruben, Laurentiu Rozylowicz and Viorel Dan Popescu. 2013. Modeling road mortality hotspots of Eastern Hermann's Tortoise in Romania. Amphibia-Reptilia 34 (2): 163-172 - get paper here
  • Jablonski, Daniel 2011. Reptiles and amphibians of Albania with new records and notes on occurrence and distribution. Acta Soc. Zool. Bohem. 75: 223–238 - get paper here
  • Kabisch, K. & Belter, H. 1968. Das Verzehren von Amphibien und Reptilien durch Vögel. Zoologische Abhandlungen aus dem Staatlichen Museum für Tierkunde in Dresden 29: 191-227
  • Kabisch, K. & Engelhard, H. 1964. Beobachtungen an Reptilien in der Umgebung von Varna-Goldsand. Aquarien Terrarien 11 (6): 194-197
  • Kantz, H. 1983. Zufallstreffer ? Nachzucht von Testudo hermanni hermanni. Sauria 5 (4): 5-6 - get paper here
  • Kantz, H. 1984. Ergänzung zum Nachzuchtbericht Testudo hermanni hermanni. Sauria 6 (4): 23-23 - get paper here
  • Kern, P. 2007. Artgerechte Haltung und Vermehrung der Griechischen Landschildkröte. Draco 8 (32): 61-73 - get paper here
  • Kirsche, W. 1967. Zur Haltung, Zucht und Ethologie der Griechischen Landschildkröte (Testudo hermanni hermanni). Salamandra 3 (1-2): 36-66 - get paper here
  • Kirsche, W. 1984. An F2-Generation of Testudo hermanni hermanni Gmelin bred in captivity with remarks on the breeding of Mediterranean Tortoises 1976-1981 Amphibia-Reptilia, 5(1): 31-35. - get paper here
  • Kirsche, W. K 1984. Bastardierung von Testudo horsfieldii (Gray) und Testudo h. hermanni Gmelin. Amphibia-Reptilia 5 (3-4): 311 - 322 - get paper here
  • Koppitz, Christian; Julian Kolleck, Melanie Kurtz, Patricia Hildebrandt & Klaus Richter 2017. Die Leier des Hermes – Einblicke in das Leben zweier Landschildkrötenarten auf der Peloponnes. Terraria-Elaphe 2017 (4): 34-43 - get paper here
  • Krizmanić, I. 2018. CONTRIBUTION TO THE HERPETOFAUNA OF SERBIA-DISTRIBUTION OF REPTILES IN KOSOVO AND METOHIJA PROVINCE. Bulletin of Natural Sciences Research 9 (2): - get paper here
  • Kuchling, G. 1981. Effect of Temperature and Photoperiod on Spermatogenesis in the Tortoise, Testudo hermanni hermanni Gmelin Amphibia-Reptilia, 2(4): 329-341. - get paper here
  • Kuchling, G. 1981. Seasonal variations in the oxygen consumption and the blood glucose concentration under low temperature conditions in the male tortoise, Testudo hermanni hermanni Gmelin Amphibia-Reptilia, 2(3): 235-241. - get paper here
  • Kwet, A. 2010. Reptilien und Amphibien Europas. Kosmos-Verlag, Stuttgart, 252 pp.
  • Kwet, Axel & Benny Trapp 2014. Liste der Reptilien Europas. Draco 15 (60): 72-79 - get paper here
  • Lapparent de Broin, F. de; Bour, R. & Perälä, J. 2006. Morphological definition of Eurotestudo (Testudinidae, Chelonii): First part. Annales de Paléontologie 92: 255–304 (erratum p. 385-386) - get paper here
  • Lapparent de Broin, F. de; Bour, R.; Parham, J.F. & Perälä, J. 2006. Eurotestudo, a new genus for the species Testudo hermanni Gmelin, 1789 (Chelonii, Testudinidae). C. R. Palevol 5: 803–811 - get paper here
  • Lescure, J.; bour, R. & Ineich, I. 2009. Jean Hermann (1738-1800), professeur d’histoire naturelle et Herpétologiste strasbourgeois. Bull. Soc. Herp. France (130-131): 1-21 - get paper here
  • Ljubisavljevic, Katarina, Georg Dzukic, Tanja D. Vukov and Milos L. Kalezic. 2014. Distribution patterns of Hermann's Tortoise Testudo hermanni Gmelin, 1789, in the region of former Yugoslavia (Testudines: Testudinidae). Herpetozoa 26 (3/4): 125-138 - get paper here
  • Ljubisavljević, Katarina; Georg Džukić, Tanja D. Vukov,Miloš L. Kalezić 2012. Morphological variability of the Hermann's tortoise (Testudo hermanni) in the Central Balkans. Acta Herpetologica 7 (2): 253-262 - get paper here
  • Loy, Anna; Ramacciato, Valeria; Gentilotti, Federica; Capula, Massimo 2007. Demography of Eurotestudo hermanni in a mesic area of Central Italy. Amphibia-Reptilia 28 (1): 87-95 - get paper here
  • Makridou, Konstantina; Charalambos Thoma, Dimitrios Bakaloudis, Christos Vlachos 2019. Sexual dimorphism of Testudo tortoises from an unstudied population in northeast Greece. Herpetology Notes 12: 229-233 - get paper here
  • Martini, S., & Schmitt, M. 2024. Additional notes on the herpetofauna of Lefkada (Ionian Islands, Greece). Natural History Sciences, 11(1) - get paper here
  • Mayer, R. 1992. Zwillinge bei Testudo hermanni hermanni Gmelin, 1789 x Testudo hermanni boettgeri Mojsisovics 1889. Sauria 14 (3): 37-38 - get paper here
  • Mazzotti, Stefano; Anna Pisapia, Mauro Fasola 2002. Activity and home range of Testudo hermanni in Northern Italy. Amphibia-Reptilia 23 (3): 305-312 - get paper here
  • Mendt,A. & Bender. C. 2001. Änderung der Bundesartenschutzverordnung in Aussicht gestellt. Elaphe 9 (1): 21-23
  • Metzingen, Gerhard Eger 2005. Gemeinsame Haltung von Testudo Hermanni Boettgeri und Testudo (hermanni) hercegovinenis - Erfahrungen eines Züchters uber einen längeren Zeitraum und Auswertung der Nachzuchtdaten. Schildkröten im Fokus 2 (2): 25-30
  • Meyer, A. 1985. Die Reptilienfauna des südlichen Peleponnes. Sauria 7 (1): 13-16 - get paper here
  • Mojsisovics, A. v. 1889. Zoogeographische Notizen über Süd-Ungarn aus den Jahren 1886— 1888. Zugleich ein III. Nachtrag zur „Fauna von Bellye und Dárda”. Mitt. naturwiss. Ver. Steiermark 25: 233-250 - get paper here
  • NAUMOV, BORISLAV; NIKOLAY TZANKOV, KRASIMIR DONCHEV, BOYAN PETROV, ANDREI STOJANOV, GEORGI POPGEORGIEV, IVELIN MOLLOV, VLADIMIR BESHKOV 2016. The Herpetofauna (Amphibia and Reptilia) of Vrachanska Planina Mountains - Species Composition, Distribution and Conservation. In: Bechev, D. & Georgiev, D. (Eds.), Faunistic diversity of Vrachanski Balkan Nature Park. ZooNotes, Supplement 3, Plovdiv University Press, Plovdiv - get paper here
  • Nekrasova, O., Tytar, V., Pupins, M., Čeirāns, A., & Skute, A. 2021. GIS Modelling of the Distribution of Terrestrial Tortoise Species: Testudo graeca and Testudo hermanni (Testudines, Testudinidae) of Eastern Europe in the Context of Climate Change. Zoodiversity, 55 (5)
  • Nemesházi, E., Bókony, V. 2023. HerpSexDet: the herpetological database of sex determination and sex reversal. Sci Data 10, 377 - get paper here
  • Neumann, A. & Wenning, J. 2004. Testudo hermanni - Schlupf in freier Natur in Deutschland. Elaphe 12 (3): 41-44
  • Nikolić, Sonja; Ana Golubović, Xavier Bonnet, Dragan Arsovski, Jean-Marie Ballouard, Rastko Ajtić, Bogoljub Sterijovski, Vuk Iković, Ana Vujović, and Ljiljana Tomović 2018. Why an Apparently Prosperous Subspecies Needs Strict Protection: The Case of Testudo hermanni boettgeri from the Central Balkans. Herp. Cons. Biol. 13 (3): 673–690 - get paper here
  • Nöllert, A.; Nöllert, C. & Ritter, A. 1986. Einige Beobachtungen zur Herpetofauna der bulgarischen Schwarzmeerküste und Südwestbulgariens (Teil 2 - Die Reptilien). Herpetofauna 8 (44): 30-34 - get paper here
  • Ortega, Zaida; Abraham Mencía and Valentín Pérez-Mellado. 2017. Notes on the thermal ecology of Testudo hermanni hermanni in Menorca (Balearic Islands, Spain). Amphibia-Reptilia 38 (1): 108 - 112 - get paper here
  • Parés-Casanova, Pere M. 2019. Adaptative directional asymmetric shape in Testudo hermanni hermanni Gmelin, 1789 (Reptilia: Testudines: Testudinidae). Herpetology Notes 12: 743-747 - get paper here
  • Parham, James F.; Macey, J. Robert; Papenfuss, Theodore J.; Feldman, Chris R.; Turkozan, Oguz; Polymeni, Rosa; Boore, Jeffrey 2006. The phylogeny of Mediterranean tortoises and their close relatives based on complete mitochondrial genome sequences from museum specimens. Molecular Phylogenetics and Evolution 38(1):50-64 [2005] - get paper here
  • Pascual, Eudald; Alba Vendrell, Laia Herrerias, Josep Marí, Albert Martínez-Silvestre & Joaquim Soler 2011. Aportacions a les variacions de pes durant el procés d’hibernació en Testudo hermanni hermanni (Gmelin, 1789). Bull. Soc. Catalana d’Herp. 19 - get paper here
  • Pawlowski, S. & Krämer, C. 2009. Bemerkungen zur Herpetologie des nordwestlichen Kroatiens. Sauria 31 (4): 19-30 - get paper here
  • Petrov B. P., N. Tzankov, H. Strijbosch, G. Popgeorgiev, V. Beshkov 2006. The herpetofauna (Amphibia and Reptilia) of the Western Rhodopes mountain (Bulgaria and Greece). In: Beron P. (ed.). Biodiversity of Bulgaria. 3. Biodiversity of Western Rhodopes (Bulgaria and Greece) I. Pensoft & Nat. Mus. Natur. Hist., Sofia, pp. 863-912
  • Pfau, B. 2019. Testudo hermanni hermanni im Albera-Gebiet in Katalonien. Teil I: der Lebensraum und das Zuchtzentrum. Elaphe 2019 (3): 80-85
  • Pfau, B. 2020. Testudo hermanni hermanni im Albera-Gebiet in Katalonien. Teil II: Zucht und Aufzucht. Elaphe 2020 (2): 74-78
  • Pfau, Beate; Albert Martínez-Silvestre & Joan Budó 2023. Testudo hermanni hermanni im Albera-Gebiet in Katalonien Teil III: Gesundheitsuntersuchungen. Elaphe 2023 (6): 74
  • Pieh, A. & Philippen, H.D. 2007. Mediterrane Landschildkröten. Draco 8 (32): 4-34 - get paper here
  • Pinya, Samuel and Miguel A. Carretero. 2011. The Balearic herpetofauna: a species update and review on the evidence. Acta Herpetologica 6 (1): 59-80 - get paper here
  • Plaitano, Annalisa & Marco A. L. Zuffi 2007. Similarities and differences in adult tortoises: a morphological approach and its implication for reproduction and mobility between species. Acta Herpetologica 2 (2): 79-86
  • Pulford, E., A. Hailey & D. Stubbs 1984. Thermal relations of Testudo hermanni in France Amphibia-Reptilia, 5(1): 37-41. - get paper here
  • Rogner, M. 2005. Griechische Landschildkröten (Testudo hermanni hermanni, T. h. boettgeri, T. h. hercegovinensis). Natur und Tier Verlag, Münster, 167 pp. - get paper here
  • Rogner, M. 2007. Griechische Landschildkröten Testudo hermanni hermanni, T. h. boettgeri, T. h. hercegovinensis. Natur und Tier Verlag, Münster, 168 pp. - get paper here
  • Rogner, M. 2012. Die Schildkröten von Son Real auf Mallorca. Reptilia (Münster) 17 (96): 64-66 - get paper here
  • Rogner, M. 2013. Die Schildkröten von Kalogria (Peloponnes). Reptilia (Münster) 18 (99): 76-78 - get paper here
  • Romano, A., Sansone, L., Cacace, A., & Biancolini, D. 2023. Diversity, distribution, habitat preferences and community assemblages of Amphibians and Reptiles in the “Cilento, Vallo di Diano e Alburni” National Park (Campania, Southern Italy). Acta Herpetologica, 18(2), 115-129 - get paper here
  • Rotat, C., D. Justet, J. Detilleux, C. Remy, L. Grolit, P. Dortu, L. Istasse and M. Diez. 2008. Morphométrie appliqué a la croissance de nouvaeau-nés et de juvéniles Testudo hermanni boettgeri Mojsisovic, 1889. Chéloniens 11:18-29
  • Roulin, Alexandre; Alia Mafli, and Kazumasa Wakamatsu 2013. Reptiles Produce Pheomelanin: Evidence in the Eastern Hermann's Tortoise (Eurotestudo boettgeri). Journal of Herpetology Jun 2013, Vol. 47, No. 2: 258-261. - get paper here
  • Sacchi, Roberto; Daniele Pellitteri-Rosa, Manuela Marchesi, Paolo Galeotti, and Mauro Fasola 2013. A Comparison among Sexual Signals in Courtship of European Tortoises. Journal of Herpetology Jun 2013, Vol. 47, No. 2: 215-221. - get paper here
  • Sacchi, Roberto; Pupin, Fabio; Rosa, Daniele Pellitteri; Fasola, Mauro 2007. Bergmann's rule and the Italian Hermann's tortoises (Testudo hermanni): latitudinal variations of size and shape. Amphibia-Reptilia 28 (1): 43-50 - get paper here
  • Sailer et al. 1997. Kurzmitteilung: Siamesische Zwillinge bei Testudo hermanni boettgeri. Herpetofauna 19 (106): 12-13 - get paper here
  • Salvi, Daniele; Pierluigi Bombi 2010. Reptiles of Sardinia: updating the knowledge on their distribution. Acta Herpetologica 5 (2): 161-178 - get paper here
  • Schaible, N. 2013. Überwinterung von Griechischen Landschildkröten (Testudo hermanni boettgeri). Reptilia (Münster) 18 (103): 66-69 - get paper here
  • Schardt, M. 2007. Die Griechische Landschildkröte (Testudo hermanni boettgeri). Natur und Tier Verlag (Münster), 64 pp. - get paper here
  • Schlüter, U. 2005. Die Herpetofauna der bulgarischen Schwarzmeerküste - Teil 1: Naturraum und Schildkröten. Elaphe 13 (3): 48-56
  • Schmidt B, Kranželić D, Majetić Đ, Lauš B, Štih A, Koren T 2020. Distribution and conservation status of the herpetofauna of Dugi Otok Island, Croatia. Herpetozoa 33: 191-205 - get paper here
  • Schmidt, Hans Ulrich 2006. Testudo hermanni hermanni on Minorca - Part 5. Radiata 15(2): 27-29.
  • Schweiger, M. 1992. Das Stachelschwein Hystrix cristata (LINNAEUS, 1758) als populationslimitierender Faktor von Testudo hermanni hermanni (GMELIN, 1789). Salamandra 28 (1): 86-88 - get paper here
  • Sehnal, P. & Schuster, A. 1999. Herpetologische Beobachtungen auf der Kvarnerinsel Cres, Kroatien. Ergebnisse von fünf Exkursionen. Herpetozoa 12 (3-4): 163-178 - get paper here
  • Shah, R. V. 1960. The mechanisms of carapacial and plastral hinges in Chelonians. Breviora (130): 1-15 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, R. and Rossi, R. 2020. First report on the herpetofauna of Meganissi (Lefkada, Ionian Islands, Greece). Herpetology Notes 13: 81-84. - get paper here
  • Sos, T. 2008. Review of recent taxonomic and nomenclatural changes in European Amphibia and Reptilia related to Romanian herpetofauna. Herpetologica Romanica 2: 61-91
  • Sos, T., Daróczi, Sz., Zeitz, R. & L. Pârâu 2008. Notes on morphological anomalies observed in specimens of Testudo hermanni boettgeri Gmelin, 1789 (Reptilia: Chelonia: Testudinidae) from Southern Dobrudja, Romania. North-Western Journal of Zoology 4 (1): 154-160 - get paper here
  • Stoev, Pavel 2000. On the distribution, biology and ecology of amphibians and reptiles in the Derventski Heights and the Sakar Mountain, South-East Bulgaria Hist. Nat. Bulgarica 12: 59–69 - get paper here
  • Strachinis, Ilias; Dimitris Aravanis 2017. Additions to the known herpetofauna of the Island of Ithaki, Ionian Sea, Greece. Herpetozoa 30 (1/2): 64 - 66 - get paper here
  • Thomas, G. 1969. Fortpflanzungsverhalten bei jungen Griechischen Landschildkröten (Testudo h. hermanni). Salamandra 5 (3-4): 147-148 - get paper here
  • Tóth, Tamás, László Krecsák, Thomas Madsen and Beáta Újvári 2002. Herpetofaunal locality records on the Greek Islands of Corfu (Amphibia, Reptilia). Herpetozoa 15 (3/4):149-169 - get paper here
  • Toth, Tamas; Miklós Heltai, Andrea Keszi, Goran Susic, Levente Moharos, Balazs Farkas, Csaba Geczy, Orsolya Torda, Janos Gal 2017. Herpetofauna inventory of the small islands of the Cres-Losinj Archipelago (North Adriatic Sea, Croatia) (Amphibia; Reptilia). Herpetozoa 30 (1/2): 21 - 28 - get paper here
  • Trapp, B. 2004. Testudo hermanni boettgeri, Griechische Landschildkröte (MIJSIOVICS 1889). Reptilia (Münster) 9 (46): 51-54 - get paper here
  • Trapp, B. 2007. Reptilien und Amphibien des griechischen Festlandes. Natur und Tier Verlag (Münster), 300 pp. - get paper here
  • Trapp, B. & Valverde, J. 2004. Testudo hermanni boettgeri. Reptilia (GB) (35): 39-42 - get paper here
  • Trutnau, L. 1975. Europäische Amphibien und Reptilien. Belser Verlag, Stuttgart 212 pp.
  • TTWG; Anders G.J. Rhodin, John B. iverson, Roger Bour, Uwe Fritz, Arthur GeoRges, H. BRadley Shaffer, and Peter Paul van Dijk 2017. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (8th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Pritchard, P.C.H., and Mittermeier, R.A. (Ed Chelonian Research Monographs 7:1–292. doi: 10.3854/crm.7.checklist.atlas.v8.2017 - get paper here
  • TTWG; Rhodin, A.G.J., Iverson, J.B., Bour, R., Fritz, U., Georges, A., Shaffer, H.B., and van Dijk, P.P. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Stanford, C.B., Goode, E.V., Buhlmann, K.A., and Mittermeier, R.A. (Eds.). Chelonian Research Monographs 8:1–472. doi:10.3854/crm.8.checklist.atlas.v9.2021. - get paper here
  • Türkozan, Oğuz; Adem Özdemir, and Ferhat Kiremit 2008. International Testudo Trade. Chelonian Conservation and Biology 7(2): 269 - get paper here
  • Turkozan, Oguz; Can Yilmaz, Şule Karakaya, Sezgin Karaman & Celal Ulger 2019. Distribution, Size, and Demographics of Eastern Hermann's Tortoise, Testudo hermanni boettgeri, in Turkey. Chelonian Conservation and Biology Dec 2019 Vol. 18, No. 2: 210-216 - get paper here
  • Türkozan, Oğuz; Ferhat Kiremit, Ertan Taşkavak, Kurtuluş Olgun 2005. Status, distribution, and population structure of land tortoises in European Thrace and northwestern Anatolia. Russ. J. Herpetol. 12 (3): 187-194. - get paper here
  • TZORAS, ELIAS; ALEXIOS PANAGIOTOPOULOS & DANIEL JABLONSKI. 2022. Outside the home: Hermann’s Tortoise, Testudo hermanni (Chelonia: Testudinidae), on Milos Island, Cyclades, Greece. Herpetology Notes 15: 625–627. - get paper here
  • van der Kuyl, Antoinette C.; Donato L. Ph. Ballasina, John T. Dekker, Jolanda Maas, Ronald E. Willemsen and Jaap Goudsmit 2002. Phylogenetic Relationships among the Species of the Genus Testudo (Testudines: Testudinidae) Inferred from Mitochondrial 12S rRNA Gene Sequences. Molecular Phylogenetics and Evolution 22: 174-183 - get paper here
  • Vecchio, Silvia Del; Russell L. Burke, Lorenzo Rugiero, Massimo Capula, and Luca Luiselli 2011. Seasonal Changes in the Diet of Testudo hermanni hermanni in Central Italy. Herpetologica 67 (3): 236-249. - get paper here
  • Vetter, H. 2006. Hermann’s Tortoise - Testudo hermanni. Edition Chimaira, Frankfurt, 320 pp.
  • Vetter, H. 2006. Griechische Landschildkröte, Dalmatinische und Italienische Landschildkröte. Testudo boettgeri, T. hercegovinensis und T. hermanni. Edition Chimaira, Frankfurt, 320 pp.
  • Vilardell-Bartino, Albert, Joan Budó, Xavier Capalleras, Joan Ferrer & Eduard Filella 2011. Dades sobre la incidència d’un incendi forestal en la població de tortuga mediterrània (Testudo hermanni hermanni) a la serra de l’Albera (Alt Empordà; Catalunya). Bull. Soc. Catalana d’Herp. 19 - get paper here
  • Vinke, T. & Vinke, S. 2004. Testudo hercegovinensis WERNER, 1899 – die Dalmatinische Landschildkröte. Schildkröten im Fokus 1 (1): 22-34
  • Vinke, Thomas; Vinke, Sabine 2006. Konkurrierende Modelle der wissenschaftlichen Systematik – oder Eurotestudo und Agrionemys contra Testudo. Schildkröten im Fokus - Sonderband 1:9-12.
  • Wermuth, Heinz 1952. Testudo hermanni robertmertensi n. subsp. und ihr Vorkommen in Spanien. Senckenbergiana 33 (1/3): 157-164
  • Wiechert, J. 2007. Ein seltener Ektoparasitenbefall: Schlangenmilben (Ophionyssus natricis) bei Griechischen Landschildkröten der Unterart Testudo hermanni boettgeri. Draco 8 (32): 47-49 - get paper here
  • Wilson, D.S. et al. 2003. Estimating age of turtles from growth rings: a critical evaluation of the technique. Herpetologica 59 (2): 178-194 - get paper here
  • Wirth, M. 2008. Testudo hermanni hermanni. Reptilia (Münster) 13 (71): 51-54 - get paper here
  • Wirth, M. 2010. Zwischen Müll und Minen: die Dalmatinische Landschildkröte Testudo hermanni hercegovinensis. Teil 2. Terraria (23) 5 (3): 50-58.
  • Wirth, M. 2010. Ein Hauch von Mittelmeer - Bepflanzung von Freilandanlagen für mediterrane Landschildkröten. Draco 11 (41): 32-43 - get paper here
  • Wirth, M. 2010. Zwischen Müll und Minen: die Dalmatinische Landschildkröte Testudo hermanni hercegovinensis. Teil 1 Terraria (22) 5 (2): 44-47.
  • Wirth, M. 2013. Das Schildkrötenjahr - Freilandbiologie und Haltung europäischer Landschildkröten über den Jahresverlauf. Natur und Tier Verlag, Münster, 279 pp. - get paper here
  • Wirth, Michael 2012. Alles der Wärme zuliebe – die Optimierung von Landschildkrötengehegen. Draco 12 (49): 62-69
  • Wirth, Michael 2012. Immer der Größe angemessen – die erfolgreiche Haltung und Aufzucht von Schlüpflingen und Jungtieren im Freilandterrarium. Draco 12 (49): 50-61
  • Wirth, Michael 2012. Die Vorbereitung der Eiablage:Auf den Legehügel kommt es an!. Draco 12 (49): 40-49
  • Wirth, Michael 2013. Vorbild Natur: Die Ernährung europäischer Landschildkröten im natürlichen Lebensraum. Draco 13 (54): 6-19 - get paper here
  • Wirth, Michael 2013. Überwinterung europäischer Landschildkröten - naturnah oder kontrolliert? Draco 14 (55): 66-73 - get paper here
  • Wirth, Michael 2013. Toskana. Ein herpetologischer Überblick. Terraria-Elaphe 2013 (3): 41-24 - get paper here
  • Wirth, Michael 2013. Heu in der Schildkrötenernährung – wertvoll und lebensnotwendig. Draco 13 (54): 54-66 - get paper here
  • Wirth, Michael 2013. Schildkrötenernährung, aber artgerecht – eine Praxisanleitung. Draco 13 (54): 20-29 - get paper here
  • Woldring, L. 2005. Stamboek Testudo hermanni hermanni (GMELIN 1798). Lacerta 63 (6): 257-260 - get paper here
  • Wütschert, R. 1984. Neues uber die Reptilienfauna der Insel Korfu. Salamandra 20 (4): 221-228 - get paper here
  • Zauner, Jochen 2023. Herpetologische Beobachtungen auf dem Peloponnes (Griechenland). Sauria 45 (4): 1-17
  • Zenboudji, Saliha; Véronique Arnal, Marc Cheylan, Albert Bertolero, Geoffrey Druelle, Marie-Pierre Dubois & Claudine Montgelard 2018. Isolation and Characterization of 32 Microsatellite Markers in Hermann's Tortoise, Testudo hermanni (Testudinidae). Chelonian Conservation and Biology Dec 2018 Vol. 17, No. 2: 291-297 - get paper here
  • ŽIVKOV, M. T. & RAIKOVA-PETROVA, G. N. & TRICHKOVA, T. A. 2009. Population biology and ontogenetic allometry of Testudo hermanni boettgeri MOJSISOVICS, 1889, in a marginal population in Bulgaria. Herpetozoa 22: - get paper here
 
External links  
Is it interesting? Share with others:

As link to this species use URL address:

https://reptile-database.reptarium.cz/species?genus=Testudo&species=hermanni

without field 'search_param'. Field 'search_param' is used for browsing search result.



Please submit feedback about this entry to the curator