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Trachylepis sulcata (PETERS, 1867)

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Higher TaxaScincidae, Mabuyinae (Mabuyini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Western Rock Skink 
SynonymEuprepes olivaceus PETERS 1862: 21 (pre-occupied)
Euprepes sulcatus PETERS 1867: 20 (replacement name for E. olivaceus)
Mabuia sulcata — WERNER 1910: 345
Trachylepis sulcata nigra (WERNER 1915)
Mabuya sulcata ansorgii — MERTENS 1938: 438 [part]
Mabuya sulcata ansorgii —HELLMICH 1957b: 55
Mabuya sulcata ansorgii —BRANCH 1998: 157
Mabuya sulcata ansorgei — LAURENt 1964: 74
Mabuya sulcata — GREER et al. 2000
Mabuya sulcata — KABISCH 2000
Euprepis sulcatus — MAUSFELD et al. 2002
Mabuya sulcata — BAUER & BRANCH 2003
Trachylepis sulcata — BAUER 2003
Mabuya sulcata — COOPER 2005
Trachylepis sulcata sulcata — BATES et al. 2014: 267
Trachylepis sulcata sulcata — BUTLER et al. 2019
Trachylepis sulcata — CERIACO et al. 2024: 106 
DistributionNamibia, Republic of South Africa (Karroid areas of the Cape and adjacent OFS), SW Angola

Type locality: Neu-Barmen [= Otjimbingue, Otjozondjupa Region, Namibia], by implication [Euprepes olivaceus].

ansorgi: Angola; Terra typica: Caeonda, Benguella, Angola.

nigra: Lüderitz Bay, SC Namibia.  
Reproductionapparently both oviparous and viviparous reproduction (Shine 1985) 
TypesSyntypes: ZMB 4209 (n = 3), 4210 (n = 3), Neu Barmen (=Gross Barmen, Namibia), collected C.H. Hahn; MNHN 1470 (1867.133, 2792a) (see Brygoo, 1985).
Syntypes: ZMB 4209, 4210, MNHN 1470 ? (fide BRYGOO 1985) [Euprepes olivaceus] 
DiagnosisDiagnosis: Ceriaco et al. 2024: 110 
CommentSynonymy: Euprepes olivaceus Peters, 1862: 21 is preoccupied by Euprepes olivaceus Gray, 1839.

Subspecies: Trachylepis sulcata ansorgii (BOULENGER 1907) has been elevated to full species status by BAUER et al. 2023.

Distribution: see map in Ceriaco et al. 2024: 110 (Fig. 91).

Phylogenetics: Mito-nuclear discordance was observed by Butler et al. 2024 with respect to the geographic boundary between T. ansorgii and T. sulcata. However, both species are supported by molecular data. 
References
  • Baptista NL, António T, Branch WR. 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2) [Special Section]: 96–130 (e203) - get paper here
  • Bates, M.F. 1995. Life History Notes - Mabuya sulcata sulcata. African Herp News (24): 23-25 - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bates, M.F.; Flemming, A.F. 1995. Geographical Distribution - Mabuya sulcata sulcata. African Herp News (23): 47-47 - get paper here
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Bauer, A.M.; Günther,R. & Klipfel,M. 1995. The herpetological contributions of Wilhelm C.H. Peters (1815-1883). SSAR Facsimile Reprints in Herpetology, 714 pp.
  • Bauer, Aaron M.; Branch, William R. & Haacke, Wulf D. 1993. The herpetofauna of the Kamanjab area and adjacent Damaraland, Namibia. Madoqua (Windhoek) 18 (2): 117-145. - get paper here
  • Boulenger, G. A. 1907. Descriptions of three new lizards and a new frog, discovered by Dr. W. J. Ansorge in Angola. Ann. Mag. Nat. Hist. (7) 19: 212—214. - get paper here
  • Branch W. R.; Haagner, G.V. 1993. The skink Mabuya ivensii: new records from Zambia and Zaire, and the status of the subspecies septemlineata Laurent 1964 and the genus Lubuya Horton. Amphibia-Reptilia 14 (2): 105-115 - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Branch, William R. 1994. Herpetofauna of the Sperrgebeit [sic] Region of Southern Namibia. Herpetological Natural History 2 (1): 1-11.
  • Broadley, D.G. 2000. A review of the genus Mabuya in southeastern Africa (Sauria: Scincidae). African Journal of Herpetology, 49 (2): 87-110 - get paper here
  • Butler, B O; Ceríaco, L M P; Marques, M P; Bandeira, S; Júlio, T; Heinicke, M P & Bauer, A M; 2019. Herpetological survey of Huíla Province, Southwest Angola, including first records of Bicuar National Park. Herpetological Review 50 (2): 225-240 - get paper here
  • Butler, B. O., Ceríaco, L. M., Jackman, T. R., & Bauer, A. M. 2024. Population genetics and phylogeography of Trachylepis sulcata (Peters, 1867) and T. ansorgii (Boulenger, 1907) in south-western Africa. African Journal of Herpetology, 1-25 - get paper here
  • Ceríaco, Luis M.P.; Mariana P. Marques, Diogo Parrinha, Arthur Tiutenko, Jeffrey L. Weinell, Brett O. Butler & Aaron M. Bauer 2024. The Trachylepis (Squamata: Scincidae) of Angola: An Integrative Taxonomic Review with the Description of Seven New Species Bulletin of the American Museum of Natural History Feb 2024 Vol. 2024, No. 465: 1-153 - get paper here
  • Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2): 181–214 (e322) - get paper here
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT, H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. African Herp News (53): 35-41 - get paper here
  • Conradie, Werner; Brian Reeves , Neil Brown, & Jan A. Venter 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa. Indago 32: 81-98 - get paper here
  • Cooper Jr., W.E. 2005. Duration of movement as a lizard foraging movement variable. Herpetologica 61 (4): 363-372 - get paper here
  • Greer, A.E., Arnold, C. & Arnold, E.N. 2000. The systematic significance of the number of presacral vertebrae in the scincid lizard genus Mabuya. Amphibia-Reptilia 21 (1): 121-126 - get paper here
  • Hellmich, W. 1957. Herpetologische Ergebnisse einer Forschungsreise in Angola. Veröff. Zool. Staatssammlung München 5: 1-91 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Hewitt, John 1911. Records of South African Lacertilia and Amphibia, Addenda and Corrigenda. Annals Transvaal Museum 3 (1): 42-55. - get paper here
  • Ineich, I. & L. CHIRIO 2001. Description d'un nouveau scincidé de Republique Centrafricaine (Lacertilia: Mabuya pendeana). Bull. Soc. zool. Fr., 125 (3): 197-204 [2000, see Fretey 2023]
  • Kabisch, K. 2000. Exkursionen am Südrand des Erongo-Gebirges. Die Aquarien- und Terrarienzeitschrift 53 (4): 28-33
  • Laurent, R. 1947. Notes sur quelques reptiles appartenant a la collection du Musée Royal d'Histoire Naturelle de belgique. Bull. Mus. Hist. nat. Belg. 23 (16): 1-12
  • Lobón-Rovira J, Vaz Pinto P, S. Becker F, Tolley KA, Measey J, Bennet B, Boon B, de Sá S, Conradie W 2022. An updated herpetofaunal species inventory of Iona National Park in southwestern Angola. Check List 18(2): 289-321 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Mausfeld,P.; Vences, M. Schmitz, A. & Veith, M. 2000. First data on the molecular phylogeography of scincid lizards of the genus Mabuya. Mol. Phylogenet. Evol. 17 (1): 11-14 - get paper here
  • Peters, Wilhem Carl Hartwig 1867. Herpetologische Notizen. Monatsber. königl. Akad. Wiss. Berlin. 1867 (Januar): 13-37 - get paper here
  • Peters,W.C.H. 1862. Übersicht einiger von dem , durch seine afrikanische Sprachforschungen, rühmlichst bekannten, Hrn. Missionär C.H. Hahn bei Neu-Barmen, im Hererolande, and der Westküste von Afrika, im 21˚ südl. Br. gesammelten Amphibien, nebst Beschreibungen der neue Monatsber. Akad. Wiss. Berlin 1862: 15-26 - get paper here
  • Portik, Daniel M. 2009. Comparative Phylogeography of Two Skink Species in Southern Africa MSc thesis, Villanova, 212 pp. - get paper here
  • Portik, Daniel M.; Aaron M. Bauer & Todd R. Jackman 2010. The phylogenetic affinities of Trachylepis sulcata nigra and the intraspecific evolution of coastal melanism in the western rock skink. African Zoology 45 (2):147-159 - get paper here
  • Schleicher, Alfred 2015. Reptilien Namibias. Namibia Scientific  Society, 276 pp.
  • Schleicher, Alfred 2020. Reptiles of Namibia. Kuiseb Publishers, Windhoek, Namibia, 271 pp.
  • Sousa, Jackson Cleiton and Costa-Campos, Carlos Eduardo 2016. Erythrolamprus taeniogaster Diet. Herpetological Review 47 (4): 682 - get paper here
  • Weinell, J. L., Branch, W. R., Colston, T. J., Jackman, T. R., Kuhn, A., Conradie, W., & Bauer, A. M. 2019. A species-level phylogeny of Trachylepis (Scincidae: Mabuyinae) provides insight into their reproductive mode evolution. Molecular Phylogenetics and Evolution - get paper here
  • Werner,F. 1910. Reptilia et Amphibia. In Schultze, L., Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Südafrika. Band IV, Systematik und Tiergeographie Vertebrata B. Denkschr. Med.-Nat. Wiss. GeselI. Jena 16: 279-370 [1910] - get paper here
 
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