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Trapelus agilis (OLIVIER, 1807)

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesTrapelus agilis isolepis (BOULENGER 1885)
Trapelus agilis agilis (OLIVIER 1807)
Trapelus agilis pakistanensis RASTEGAR-POUYANI 1999
Trapelus agilis khuzistanensis RASTEGAR-POUYANI 1999 
Common NamesE: Brilliant ground agama
G: Steppenagame
Russian: прыткая агама 
SynonymAgama agilis OLIVIER 1807: 394
Agama agilis — DUMÉRIL & BIBRON 1837: 496
Agama agilis — ANDERSON 1872
Agama isolepis BOULENGER 1885: 342
Agama agilis — BOULENGER 1885: 341
Agama isolepis — BOULENGER 1887: 407
Agama kirmanensis NIKOLSKY 1899: 389
Agama kirmanensis var. brevicauda NIKOLSKY 1907: 272
Agama agilis — SMITH 1935: 221
Agama agilis agilis — WETTSTEIN & LÖFFLER 1951
Agama agilis — HAAS 1957: 66
Agama agilis — LEVITON 1959: 449
Agama kirmanensis — WERMUTH 1967: 16
Agama agilis — NILSON & ANDREN 1981: 134
Trapelus agilis — MACEY et al. 2000
Trapelus agilis — SINDACO & JEREMČENKO 2008
Laudakia kirmanensis — PYRON & BURBRINK 2013
Trapelus agilis — SHAHAMAT et al. 2020

Trapelus agilis isolepis (BOULENGER 1885)
Agama isolepis BOULENGER 1885: 342
Agama isolepis — ALCOCK & FINN 1897: 555
Agama isolepis — PROCTER 1923: 123
Agama agilis isolepis — WETTSTEIN & LÖFFLER 1951
Agama isolepis — SHAHAMAT et al. 2020

Trapelus agilis khuzistanensis RASTEGAR-POUYANI 1999
Trapelus agilis khuzistanensis RASTEGAR-POUYANI 1999
Trapelus khuzestanensis — SHAHAMAT et al. 2020 (in error)

Trapelus agilis pakistanensis RASTEGAR-POUYANI 1999
Trapelus agilis pakistanensis — KHAN 2003
Trapelus agilis pakistanensis — ADIL et al. 2020 
DistributionIran, Pakistan, India (Rajasthan, Gujarat), Russia ?, Turkmenistan, Tajikistan, Uzbekistan, Kazakhsan, China,
Iraq, Afghanistan

agilis: C Iranian Plateau, S Afghanistan, and SW Pakistan; Type locality: vicinity of Baghdad, Iraq. Changed to 110 km southeast of Isfahan city, Isfahan Province, central Iranian Plateau by Rastegar-Pouyani, 1999. This change was declared as invalid by Ananjeva et al. 2013.

isolepis: Afghanistan, Iran; Type locality: “between Magas and Bampur, Iran"

khuzistanensis: SW Iran; Type locality: SW Iran, Khuzistan Province, 5 km northwest of Haft-Gel on the road to Shushtar

pakistanensis: SE Pakistan and adjacent NW India. Type locality: Gaj-River, Kirthar Range, southeastern Pakistan

kirmanensis: Iran; Type locality: Kurin in Kirmano in Persia orientali [= Kurin, Sargad Region, Sistan and Baluchestan Province, Iran, 28.88 N 60.36 E]

brevicauda: Iran (Esfahan); Type locality: Kochrud in provincia Irak-Adschemi “Kochrud in prov. Irak-Ad-schemi” [= Kohrud or Ghohrud, Isfahan province, Iran, 33°38’ N 51°27’ E; given as 33.67 N 51.41 E by Ananjeva et al. 2020].  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 5708, and MNHN-RA 1994.1178 (formerly MNHN-RA 5708A) (2 syntypes)
Holotype: GNM 5424 (given as GNHM, Göteborg [khuzistanensis]
Holotype: SMF 63258 [pakistanensis]
Holotype: ZISP 9321, Leg: N. A. Zarudny, 21.VI.1898 [kirmanensis]
Holotype: ZISP 10335, Leg: N. A. Zarudny, 26.IV.1904 [kirmanensis var. brevicauda]
Lectotype: BMNH 74.11.23.113 [isolepis] 
DiagnosisDiagnosis (agilis): Dorsal scales subequal, homogeneous, large scales of back grading into progressively smaller scales of flanks, no distinctly larger scales among them (but see remarks below); 1-3 rows ofcallose scales anterior to vent, more prominent in males than in females.


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CommentHaas 1957 cites OLIVER 1804 as author and year.

Synonymy: partly after RASTEGAR-POUYANI 2005; Rastegar-Pouyani 1999 synonymized T. (a.) isolepis with T. agilis, but later treated T. isolepis as valid (e.g. in Shahamat et al. 2020).

Subspecies: T. a. sanguinolentus is treated here as a valid species.

Phylogenetics: Shahamat et al. 2020 found evidence for 9 species (or at least genetically different populations) of Trapelus on the Iranian plateau.

Distribution: See map in SMID et al. 2014 for distribution in Iran. In Russia fide Sindaco et al. 2008: map 100, but not in Russia fide Dunaev & Orlova 2017. The species in Russia is T. sanguinolentus fide Doronin (pers. comm., 21. June 2018).

Reference images: see Uetz et al. 2024 for high-resolution reference images for this species. 
References
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