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Trimeresurus whitteni VOGEL, DAVID & SIDIK, 2022

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Siberut Pitviper
Indonesian: Ular Mati Ekor
Siberut: Ulou Bopai pai
French: Trimérésure de Siberut
German: Siberut Grubenotter 
SynonymTrimeresurus whitteni VOGEL, DAVID & SIDIK 2022: 70
Trimeresurus hageni — SANDERS et al. 2002: 109 [partim]
Trimeresurus hageni — SANDERS et al. 2004: 729 [partim]
Trimeresurus (Trimeresurus) whitteni— MIRZA et al. 2023 
DistributionIndonesia (Siberut Island, West Sumatra Province)

Type locality: Saibi Samukop, Siberut Island, Kepulauan Mentawai, Sumatera Barat Province, Indonesia.  
Reproduction 
TypesHolotype. MZB.OPHI.2310, adult male, collected by Irvan Sidik and M. Toha in June 1994.
Paratypes (6 specimens). CAS SUR 8318, BMNH 1977.1237, RMNH 5517 (all males), BMNH 1979.267–269 (females), all from Siberut Island. 
DiagnosisDiagnosis: A large species of pitviper of the genus Trimeresurus, characterized by the combination of (1) body elongate, head long and massive in adults; (2) an overall green coloration, with irregular areas interstitial skin black and many scales irregularly edged with black producing faint, poorly contrasted crossbands; (3) a white or white and black ventrolateral stripe; (4) a dark postocular streak in most specimens, black and black and red; (5) 21 DSR at midbody; (6) 1st supralabial totally separated from nasal scale; (7) large internasals, always in contact in all examined specimens; (8) 10–11 supralabials, 4th or 4th–5th SL in contact with subocular; (9) supraoculars large but elongate, separated by 7–8 cephalic scales; (10) tail long, with a ratio TaL/TL between 0.191 and 0.209 in males and 0.144 and 0.151 in females; (11) 178–191 VEN; (12) 63–77 SC (males: 74– 77; females: 63–67); (13) eye copper to greenish copper in life; (14) cephalic scales not edged with black; (15) a red or grey postocular streak; (16) venter green, uniform; and (17) tail green as the body anteriorly, with greyish red, salmon, rusty brown or reddish brown blotches, progressively fused posteriorly making the tail entirely red or salmon. (Vogel et al. 2022)


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CommentDistribution: The species might also live on Sipora Island, just south of Siberut, from where Dring et al. (1990) reported a pitviper under the name T. sumatranus. See Vogel et al. 2022: 78 (Fig. 13) for a map. 
EtymologyNamed after Mr. Tony Whitten (10 April 1953– 29 November 2017), a leading conservationist and naturalist in Indonesia. 
References
  • Mirza ZA, H. T. Lalremsanga, Bhosale H, Gowande G, Patel H, Idiatullina SS, Poyarkov NA 2023. Systematics of Trimeresurus popeiorum Smith, 1937 with a revised molecular phylogeny of Asian pitvipers of the genus Trimeresurus Lacépède, 1804 sensu lato. Evolutionary Systematics 7(1): 91-104 - get paper here
  • Sanders, K. L.; Malhotra, A.; Thorpe, R. S. 2004. Ecological diversification in a group of Indomalayan pitvipers (Trimeresurus): convergence in taxonomically important traits has implications for species identification. Journal of Evolutionary Biology 17(4):721-731. - get paper here
  • Sanders, K.L., A. Malhotra, and R.S. Thorpe 2002. A contribution to the systematics of two commonly confused pitvipers from the Sunda Region: Trimeresurus hageni and T. sumatranus. Bulletin of the Natural History Museum London, 68(2): 107–111
  • Vogel, Gernot; Patrick David & Irvan Sidik 2022. A REVIEW OF THE COMPLEX OF Trimeresurus hageni (LIDTH DE JEUDE, 1886) (SQUAMATA: VIPERIDAE) WITH DESCRIPTIONS OF THREE NEW INSULAR SPECIES FROM INDONESIA TAPROBANICA 11 (2): 54-83
 
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