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Trimeresurus calamitas VOGEL, DAVID & SIDIK, 2022

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymTrimeresurus calamitas VOGEL, DAVID & SIDIK 2022: 62
Trimeresurus erythrurus var. ? — MÜLLER 1887: 280
Bothrops formosus — MODIGLIANI 1889: 121
Trimeresurus hageni — SANDERS et al. 2002: 109 [partim]
Trimeresurus hageni — SANDERS et al. 2004: 729 [partim]
Trimeresurus (Trimeresurus) calamitas— MIRZA et al. 2023 
DistributionIndonesia (Nias Island, North Sumatra Province)

Type locality: Nias Island  
Reproduction 
TypesHolotype. NMW 28157:5, adult female. Don. F. Steindachner, 1901, purchased from “Raff.” Paratypes (6 specimens). NHMB 9179 (male) “South Nias”, ZMH R06935 (male), ZFMK 32509 (female), BMNH 84.1.8.46 (female); MZB.OPHI 445 (female); ZMB 52039 (female) all from “Nias” without locality except MZB.OPHI 445, from Idano Gawo, Nias.
Other material (examined by Vogel et al. 2022: 5 males and 18 females). BMNH 84.12.31.14 (male); BMNH 84.1.8.47 (female); BMNH 84.12.31.13 (female); MNHN 5794, female; NMW 28155:2; NMW 28156:2; NMW 28157:4 (all males); NMW 28156:1, 3–4; NMW 28157:1–3; NMW 28159:4; NMW 28160:1–4 (all females); ZFMK 32509–10 (females); ZMB 65721 (male); ZMB 52039–41 (all females), all from Nias except MNHN 5794, with merely ―Sumatra‖ as locality, and NMW 28159:2, from “Padang”, and NMW 28159:4, which is said to come from “Deli, Sumatra”, now Medan, North Sumatra Province, both most probably in error. 
DiagnosisDiagnosis: A large species of pitviper of the genus Trimeresurus, characterized by the combination of (1) body elongate, head long and massive in adults; (2) dorsal surfaces deep or dark green, or dark bluish-green, with dorsal scales largely edged with black but non forming a pattern of crossbands; (3) 21 DSR at midbody; (4) 1st supralabial totally separated from nasal scale; (5) large internasals, always in contact; (6) usually 2 supralabials, 3rd–4th SL in contact with subocular (rarely 1st or 3rd SL only); (7) supraoculars large but elongate, separated by 5– 7 cephalic scales; (8) tail long, with a ratio TaL/TL between 0.189 and 0.208 in males and 0.143 and 0.166 in females; (9) 174–192 VEN; (10) 61–77 SC (males: 75–77; females: 61–71); (11) eye colour greenish-gold in preservative (unknown in life); (12) cephalic scales strongly and broadly edged with black but not forming streaks; (13) no postocular streak; (15) no ventrolateral stripe or, rarely, a poorly defined stripe due to a cream or pale greenish-yellow spot on each scale of the 1st DSR; (16) tail dark green as the body, with scales edged with black, marked with several, large rusty brown or reddish-brown blotches (pinkish-salmon in preservative), narrowly separated with dark green, becoming fused on the last quarter of the tail, which is entirely of the same colour as the blotches; and (17) venter pale yellowish green or pale bluish green; each ventral narrowly edged with black, especially on their central part, sometimes on the whole of their posterior margin. (Vogel et al. 2022) 
CommentDistribution: See Vogel et al. 2022: 78 (Fig. 13) for a map. 
References
  • Mirza ZA, H. T. Lalremsanga, Bhosale H, Gowande G, Patel H, Idiatullina SS, Poyarkov NA 2023. Systematics of Trimeresurus popeiorum Smith, 1937 with a revised molecular phylogeny of Asian pitvipers of the genus Trimeresurus Lacépède, 1804 sensu lato. Evolutionary Systematics 7(1): 91-104 - get paper here
  • Modigliani,E. 1889. Materiali per la fauna erpetologica dell isola Nias. Ann. Mus. civ. stor. nat. Genova 7: 113-124 - get paper here
  • Müller,F. 1887. Fünfter Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums. Verh. Naturf. Ges. Basel 8: 249-296 - get paper here
  • Sanders, K. L.; Malhotra, A.; Thorpe, R. S. 2004. Ecological diversification in a group of Indomalayan pitvipers (Trimeresurus): convergence in taxonomically important traits has implications for species identification. Journal of Evolutionary Biology 17(4):721-731. - get paper here
  • Sanders, K.L., A. Malhotra, and R.S. Thorpe 2002. A contribution to the systematics of two commonly confused pitvipers from the Sunda Region: Trimeresurus hageni and T. sumatranus. Bulletin of the Natural History Museum London, 68(2): 107–111
  • Vogel, Gernot; Patrick David & Irvan Sidik 2022. A REVIEW OF THE COMPLEX OF Trimeresurus hageni (LIDTH DE JEUDE, 1886) (SQUAMATA: VIPERIDAE) WITH DESCRIPTIONS OF THREE NEW INSULAR SPECIES FROM INDONESIA TAPROBANICA 11 (2): 54-83
 
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