Trimorphodon quadruplex SMITH, 1941

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Higher Taxa	Colubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names
Synonym	Trimorphodon biscutatus quadruplex SMITH 1941: 157 Trimorphodon quadruplex — SMITH 1942: 206 Trimorphodon biscutatus quadruplex — MERTENS 1952: 76 Trimorphodon biscutatus quadruplex — SCOTT & MCDIARMID 1984 Trimorphodon quadruplex — DEVITT et al. 2008 Trimorphodon quadruplex — WALLACH et al. 2014: 745 Trimorphodon quadruplex — MARTÍNEZ-FONSECA et al. 2024
Distribution	Nicaragua, Honduras, SE Guatemala, El Salvador, Costa Rica Type locality: “Esteli, Nicaragua”
Reproduction	oviparous
Types	Holotype: USNM 89476, female
Diagnosis	Additional details, e.g. a detailed description or comparisons (376 characters), are available for collaborators and contributors. Please contact us for details.
Comment	Habitat: partly arboreal (Harrington et al. 2018).
Etymology	Presumably named after the color pattern wherein each primary blotch is completely divided into four secondary blotches by the presence of pale centers, hence “quadruplex” (four-fold weave)
References	Devitt, Thomas J.; Travis J. LaDuc, and Jimmy A. McGuire 2008. The Trimorphodon biscutatus (Squamata: Colubridae) Species Complex Revisited: A Multivariate Statistical Analysis of Geographic Variation. Copeia 2008 (2): 370–387 - get paper here Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here Jadin, Robert C., and Eric N. Smith. 2010. The hemipenis of Trimorphodon quadruplex. Herpetological Review 41(2): 157-159 - get paper here Lillywhite, H.B. 2022. Discovering snakes in wild places. ECO Publishing, Rodeo, NM, 164 pp. - get paper here Martínez-Fonseca JG, Holmes IA, Sunyer J, Westeen EP, Grundler MR, Cerda PA, Fernández-Mena MA, Loza-Molina JC, Monagan IV Jr., Nondorf D, Pandelis GG, Rabosky ARD 2023. A collection and analysis of amphibians and reptiles from Nicaragua with new country and departmental records. Check List 20(1): 58-125 - get paper here McCranie, James R. 2014. Geographic Distribution: Trimorphodon quadruplex (Central American lyresnake; colcuate). Herpetological Review 45 (2): 286 - get paper here McCranie, James R. 2015. A checklist of the amphibians and reptiles of Honduras, with additions, comments on taxonomy, some recent taxonomic decisions, and areas of further studies needed. Zootaxa 3931 (3): 352–386 - get paper here McCranie, James R., and Alexander Gutsche 2016. The herpetofauna of islands in the Golfo de Fonseca and adjacent waters, Honduras. Mesoamerican Herpetology 3 (4): 842–899 - get paper here Mertens, R. 1952. Die Amphibien und Reptilien von El Salvador. Abh. senckenb. naturf. Ges. (Frankfurt) (No. 487): 120 pp. Scott, N. J., & MCDIARMID, R. W. 1984. Trimorphodon biscutatus. Catalogue of American Amphibians and Reptiles (353): 1-4. - get paper here Smith, Hobart M. 1941. Notes on the snake genus Trimorphodon. Proc. US Natl. Mus. 91 (3130): 149-168 - get paper here Solís, J. M., L. D. Wilson, and J. H. Townsend. 2014. An updated list of the amphibians and reptiles of Honduras, with comments on their nomenclature. Mesoamerican Herpetology 1: 123–144 - get paper here Sunyer, Javier 2014. An updated checklist of the amphibians and reptiles of Nicaragua. Mesoamerican Herpetology 1 (2): 186–202. - get paper here Sunyer, Javier & José́ Gabriel Martínez-Fonseca 2023. An updated country checklist to the amphibians and reptiles of Nicaragua. REVISTA NICARAGÜENSE DE BIODIVERSIDAD (100): 1-25 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
External links	IUCN Red List National Center for Biotechnology Information http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1643%2FCH-07… Google images

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