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Tropidodipsas tricolor GRÜNWALD, TORIBIO-JIMÉNEZ, MONTAÑO-RUVALCABA, FRANZ-CHÁVEZ, PEÑALOZA-MONTAÑO, BARRERA-NAVA, JONES, RODRIGUEZ, HUGHES, STRICKLAND & REYES-VELASCO, 2021

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Tricolor Snailsucker
S: Caracolera tricolor 
SynonymTropidodipsas tricolor GRÜNWALD, TORIBIO-JIMÉNEZ, MONTAÑO-RUVALCABA, FRANZ-CHÁVEZ, PEÑALOZA-MONTAÑO, BARRERA-NAVA, JONES, RODRIGUEZ, HUGHES, STRICKLAND & REYES-VELASCO 2021: 238 
DistributionMexico (Guerrero)

Type locality: 1.5 km east of Río Verde, Municipio de Atoyac de Álvarez, (17.3131°, -100.1969°, datum=WGS84, 971 m a.s.l.), Guerrero, Mexico (Fig. 3A)  
Reproduction 
TypesHolotype. INIRENA 2800 (original field num- ber CIG 1837). Juvenile male, collected on August 14th, 2020 by Mi- guel A. Peñaloza-Montaño, Jason M. Jones and Jacobo Reyes-Velasco (Fig. 1 in Grünwald et al. 2021).
Paratypes. (2) (Figs. 2C–D) INIRENA 2799 (original field number CIG 1596). Adult of unknown sex, DOR, collected at 26 km N of Putla Villa de Guerrero, on Putla Villa de Guerrero - Oaxaca Hwy., Municipality of Putla de Guerrero (17.1494°, -97.8709°, datum=WGS84, 1785 m a.s.l.), Oaxaca, Mexico, on September 4th, 2019 by Chris- toph I. Grünwald, Christopher M. Rodriguez and Carmen Mendoza-Portilla; INIRENA 2798 (original field number CIG 1863). Adult female, DOR, collected at 4.5 km NW of Mixtecapa, on the road to Malinaltepec, Municipio de Malinaltepec, (17.2539°, -98.6406°, datum=WGS84, 1815 m a.s.l.), Guerrero, Mexico on October 21st, 2020 by Tziuhtécatl Sánchez-Luna. 
DiagnosisDiagnosis: “Tropidodipsas tricolor sp. nov. is placed in the genus Tropidodipsas based on phylogenetic evidence (Fig. 6). It belongs to the Tropidodipsas fasciata group as defined by Kofron (1987) based on possessing a laterally compressed body, head distinctly wider than neck, protruding eyes of moderate to large diameter, vertebral and paravertebral scales not wider than other dorsal scales, 17 maxillary teeth, 18 dentary teeth, postmental scale absent. The species differs from all described species of Tropidodipsas based on the following combination of characters: (1) 15/15/15 rows of smooth dorsal scales with no enlarged vertebral row; (2) prefrontal broader than long, entering the orbit; (3) loreal square, equally long as high, not entering the orbit; (4) 183 ventral scales in males, 183 in females; (5) 78–79 divided subcaudals in males and 79 in females; (6) 19–22 reddish orange, light-edged bands on body, most with dark brown or black medial stippling, giving the snake a “tricolor” effect; 8 pale bands on tail; (7) ground color black or nearly so; (8) iris chocolate brown; (9) TL/SVL 0.31 in one male specimen, and 0.30 in one female specimen.” (Grünwald et al. 2021: 238)


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CommentDistribution: See Grünwald et al. 2021: 252 (Fig. 10) for a map. 
EtymologyNamed after the tricolor pattern of the black dorsal coloration interspersed by light dorsal bands of cream and reddish orange. 
References
  • Grünwald CI, Toribio-Jiménez S, Montaño-Ruvalcaba C, Franz-Chávez H, Peñaloza-Montaño MA, Barrera-Nava EY, Jones JM, Rodriguez CM, Hughes IM, Strickland JL, Reyes-Velasco J 2021. Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species. Herpetozoa 34: 233-257 - get paper here
 
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