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Tropidophorus grayi GÜNTHER, 1861

IUCN Red List - Tropidophorus grayi - Least Concern, LC

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Common NamesE: Gray's Keeled Skink 
SynonymTropidophorus grayi GÜNTHER 1861: 189
Tropidophorus grayi — GÜNTHER 1873: 165
Tropidophorus grayi — BOULENGER 1887: 364
Tropidophorus concincinensis GRAY 1845: 101 (not DUMÉRIL & BIBRON)
Enoplosaurus insignis SAUVAGE 1879: 211
Tropidophorus Grayi — FISCHER 1884: 43
Tropidophorus grayi — DE ROOIJ 1915: 279
Tropidophorus grayi — TAYLOR 1922
Tropidophorus grayi — BROWN & ALCALA 1980: 222
Tropidophorus grayi— GREER & BISWAS 2004
Tropidophorus aphilus — UNKNOWN AUTHOR (nom. nud.)
Tropidophorus apulus — HIKIDA et al. 2003 (nom. nud.) 
DistributionPhilippines (Panay, Luzon, Polillo,, Leyte, Negros, Masbate, Cebu)

Type locality: Philippines.  
TypesSyntypes: BMNH 1946.8.20.86-88 (formerly xv.78a-e), Philippines, collected H. Cuming. 
DiagnosisDEFINITION. A relatively large Tropidophorus, snout-vent length 83.8-113.2 mm for 10 mature males and 85.5-119.3 mm for 13 mature females; snout strongly tapered, round-pointed, its length 32-35 % of head length and 44-52 % of head breadth, head breaiith 73 - 75% of head length and 15 - 19% of snout vent length; eye relatively large, its diameter 75 - 93% of snout length and 27 - 31% of head breadth; rostral much broader than high, broadly concave dorsally; a pair of fronto-nasals; anterior loreal about as broad as or broader than long, about the same size as posterior loreal; prefrontals separated completely or only anteriorly by an azygous shield; frontal long and narrow, in contact with 2 or 3 supraoculars, much longer than paired frontoparietals; 4 large supraoculars; interparietal large, shaped like a wide shield; parietals large, separated hv interparietal and 2 or 3 small shields posteriorly; no nuchals: 6 or 7 upper labials, 5th beneath center of eye; 5 or 6 lower labials; head shields with prominent keels or ridges; dorsal and lateral scales with strong to moderate keels, those of tail ending in sharp spines; 24 - 28 midbody scale rows; 30 - 34 transverse rows along vertebral line between parietals and base of tail; preanals enlarged; limbs well developed, length of extended hind limb 76 - 95% of axilla-groin distance and 38 - 46% of snout-vent length; 16 - 20 lamellae beneath 4th toe; 5 - 7 lamellae beneath 1st toe; rank of toes longest to shortest 4, 3, 2. 5, 1 (or 2 and 5 nearly equal). (Brown & Alcala 1980: 223). 
CommentSynonymy mainly after BROWN & ALCALA 1980. Populations of Tropidophorus from Sulawesi (Indonesia) have been referred to T. grayi, but Hikida et al. (2003) described them as a new species, T. baconi.

Tropidophorus aphilus and T. apulus are nomina nuda according to HIKIDA et al. (2003) and the latter may be a misspelling of the former. Both appear to refer to T. grayi and are thus listed here. 
EtymologyNamed after British zoologist John Edward Gray (1800 – 1875), Keeper of Zoology at the British Museum in London from 1840 until 1874, although he started working at the museum in 1824. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Binaday, Jake Wilson B.; Ace Kevin S. Amarga, Ernesto S. Barrameda Jr., and Bobet Jan M. Bonagua 2017. Amphibians and Reptiles in the Vicinity of Bulusan Lake, Bulusan Volcano Natural Park, Sorsogon, Philippines. Philippine Journal of Science 146 (3): 339-351
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Brown, WC. & A.C. ALCALA 1980. Philippine Lizards of the family Scincidae. Silliman Univ. Nat. Sci., Dumaguete City, Mon., Ser. 2: i-xi + 1-246.
  • Decena, S. C. P., Macasait Jr, D. R., & Arguelles, M. S. 2023. Species Accounts, Assemblage, and Microhabitats of Amphibians and Reptiles of Northeastern Leyte, Philippines. Philippine Journal of Science, 152(1), 1-34
  • Ferner, John W., Rafe M. Brown, Rogelio V. Sison and Robert S. Kennedy 2000. The amphibians and reptiles of Panay Island, Philippines. Asiatic Herpetological Research 9: 1-37 - get paper here
  • Fischer, J. G. 1884. Herpetologische Bemerkungen. Abh. Nat. Ver. Hamburg 8 (2): 43-51 (3-11)
  • Gaulke, M. 2011. The herpetofauna of Panay Island, Philippines. Edition Chimaira, 390 pp.
  • Gaulke, M. 2012. Grays Kielskink (Tropidophorus grayi GÜNTHER 1861). Terraria-Elaphe 2012 (4): 76-81 - get paper here
  • Gojo-Cruz, Paul Henric P. and Leticia E. Afuang 2018. The Zoogeographic Significance of Caraballo Mountain Range, Luzon Island, Philippines With Focus on the Biogeography of Luzon’s Herpetofauna. Philippine Journal of Science 147 (3): 393-409 - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • Greer, A.E. & Biswas, S. 2004. A Generic Diagnosis for the Southeast Asian Scincid Lizard Genus Tropidophorus Dumeéril and Bibron, 1839 with Some Additional Comments on Its Morphology and Distribution. Journal of Herpetology 38 (3): 426-430 - get paper here
  • Günther, A. 1861. Second list of Siamese reptiles. Proc. Zool. Soc. London 1861: 187-189 - get paper here
  • Günther, A. 1861. Second list of Siamese reptiles. Ann. Mag. Nat. Hist. (3) 8: 266-268 - get paper here
  • Günther,A. 1873. Notes on some reptiles and batrachians obtained by Dr. Bernhard Meyer in Celebes and the Philippine Islands. Proc. Zool. Soc. London 1873: 165-172 - get paper here
  • Harbig, P. 1999. Zur Terrarienhaltung und Zucht von Tropidophorus grayi Günther, 1861. Sauria 21 (3): 35-42 - get paper here
  • Hikida, T.; Riyanto, A. & Ota, H. 2003. A new water skink of the genus Tropidophorus (Lacertilia: Scincidae) from Sulawesi, Indonesia. Current Herpetology 22 (1): 29-36 - get paper here
  • Klemmer K. & GAULKE M. 1993. Zur Kenntnis von Brachymeles und Tropidophorus (Sauria: Scincidae). Salamandra 29 (1): 55-64 - get paper here
  • Pianka, E.R. & Vitt, L.J. 2003. Lizards - Windows to the Evolution of Diversity. University of California Press, Berkeley, 347 pp. [review in Copeia 2004: 955] - get paper here
  • Sauvage, H. E. 1879. Sur un Chalcidien de genre nouveau, voisin du Tribolonote. Bull. Soc. philom. Paris (7) 3: 210-212 - get paper here
  • SUPSUP, Christian E.; Nevong M. PUNA, Augusto A. ASIS, Bernard R. REDOBLADO, Maria Fatima G. PANAGUINIT, Faith M. GUINTO, Edmund B. RICO, Arvin C. DIESMOS, Rafe M. BROWN and Neil Aldrin D. MALLARI 2016. Amphibians and Reptiles of Cebu, Philippines: The Poorly Understood Herpetofauna of an Island with Very Little Remaining Natural Habitat. Asian Herpetological Research 2016, 7(3): 151–179 DOI: 10.16373/j.cnki.ahr.150049 - get paper here
  • Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
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