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Tropiocolotes steudneri (PETERS, 1869)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Algerian Sand Gecko
G: Steudners Zwerggecko 
SynonymGymnodactylus steudneri PETERS 1869
Stenodactylus petersii BOULENGER 1885: 18
Gymnodactylus steudneri — BOULENGER 1885: 34
Stenodactylus steudneri — BOULENGER 1887
Tropiocolotes steudneri — BOULENGER 1891
Tropiocolotes nattereri STEINDACHNER 1901
Tropiocolotes steudneri — LOVERIDGE 1947: 52
Tropiocolotes steudneri — WERMUTH 1965: 184
Tropiocolotes steudneri — KLUGE 1993
Tropiocolotes (Tropiocolotes) steudneri — RÖSLER 1995: 160
Tropiocolotes steudneri — SCHLEICH, KÄSTLE & KABISCH 1996: 280
Tropiocolotes cf. steudneri — ANDERSON 1999
Tropiocolotes cf. steudneri — RASTEGAR-POUYANI et al. 2008
Tropiocolotes steudneri — TRAPE, CHIRIO & TRAPE 2012
Tropiocolotes steudneri — MACHADO et al. 2020 
DistributionSE Libya, Egypt, Sinai,
N Sudan (Jumhūriyyat), Ethiopia, Jordan, Iran ?

Type locality: Sennar [Sudan]  
Reproductionoviparous 
TypesHolotype: lost, was ZMB 5476 (lost fide BAUER et al. 1995) 
DiagnosisDiagnosis: Dorsal scales homogeneous, imbricate, weakly keeled (under magnification) 51 around middle of body; subdigitallamellae distinctly tricarinate; first pair of chin shields (postmentals) in broad contact, second pair small, separated by three scales; hind limb reaches nearly to axilla. (Diagnosis based on Iranian specimens only, fide Anderson 1999: 196). 
CommentSimilar species: Tropiocolotes naybandensis, T. steudneri, T. nattereri. Pictures of T. steudneri at http://herpetology.com/1phs31.html are in fact T. nattereri (Pierre Andre Crochet, pers. comm.).

Distribution: T. steudneri appears not to occur in Israel. All records up to know are based on misidentification of T. nattereri. SINDACO & JEREMČENKO 2008 (map 78) show one (dubious ?) record from Iran. Phylogenetic results of Machado et al. (2021) grouped Tropiocolotes steudneri from Niger and southern Algeria (Tassili n’Ajjer and Ahaggar) within Tropiocolotes nubicus Baha El Din, 1999, while T. steudneri specimens were only found east of the Nile River (fide Rachid Rouag, pers. comm., 8 May 2024). See map in Machado et al. 2020 (Fig. 1). 
EtymologyNamed after Hermann Steudner (1832-1863), naturalist and explorer. 
References
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Arnold, E. N. 1977. Little-known geckoes (Reptilia: Gekkonidae) from Arabia with descriptions of two new species from the Sultanate of Oman. The Scientific Results of the Oman Flora and Fauna Survey, 1975. Journal of Oman Studies, Special Report, (1):81—110 [1975]
  • Arnold, U. 2006. Meine “persönliche” Entdeckung von Tropiocolotes steudneri (PETERS 1869) mit Anmerkungen zu seinem Verhalten im Freiland und Terrarium. Sauria 28 (3): 23-31 - get paper here
  • Baha el Din, S. 2006. A guide to the reptiles and amphibians of Egypt. The American University in Cairo Press, Cairo. xvi + 359 pp.
  • Baha EL DIN, S. M. 1999. A new species of Tropiocolotes (Reptilia: Gekkonidae) from Egypt. Zoology in the Middle East 19: 17-26. - get paper here
  • Bauer, A.M.; Günther,R. & Klipfel,M. 1995. The herpetological contributions of Wilhelm C.H. Peters (1815-1883). SSAR Facsimile Reprints in Herpetology, 714 pp.
  • Bauer, Aaron M.; Jonathan C. DeBoer , Dylan J. Taylor 2017. Atlas of the Reptiles of Libya. Proc. Cal. Acad. Sci. 64 (8): 155-318 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Gholamifard, Ali; Ali Gholamhosseini, Nasrullah Rastegar-Pouyani,Hamid Reza Esmaeili and Haji-Gholi Kami 2010. First Records of Tropiocolotes steudneri Peters, 1869 and Hemidactylus flaviviridis Rüppell, 1840 (Sauria: Gekkonidae) from Fars Province, Iran. Asian Herpetological Research 1 (1): 61-63 - get paper here
  • Ibrahim, Adel A. 2013. The Herpetology of the Suez Canal Zone, Egypt. Vertebrate Zoology 63 (1): 87–110 - get paper here
  • KRAUSE, VALENTINA; FARAHAM AHMADZADEH, MOSTAFA MOAZENI, PHILIPP WAGNER & THOMAS M. WILMS 2013. A new species of the genus Tropiocolotes Peters, 1880 from western Iran (Squamata: Sauria: Gekkonidae). Zootaxa 3716 (1): 022–038 - get paper here
  • Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • Machado, Luís, Daniele Salvi, D James Harris, José C Brito, Pierre-André Crochet, Philippe Geniez, Faraham Ahmadzadeh, and Salvador Carranza. 2020. Systematics, Biogeography and Evolution of the Saharo-Arabian Naked-Toed Geckos Genus Tropiocolotes. Molecular Phylogenetics and Evolution 155: 106969 [print 2021] - get paper here
  • Milto, Konstantin D. 2017. New Records of Reptiles on the Red Sea Coast, Egypt, with Notes on Zoogeography. Russ. J. Herpetol. 24 (1): 11-21 - get paper here
  • Milto, Konstantin D.; Samy A. Saber, Abdullah M. Nagy, Roman A. Nazarov, Daniel A. Melnikov, Natalia B. Ananjeva 2019. First Report on the Reptile Diversity of Wadi El Gemal National Park, Eastern Desert, Egypt. Russ. J. Herpetol. 26 (3): 175-184 - get paper here
  • Peters, WILHELM C. H. 1869. Eine Mittheilung über neue Gattungen und Arten von Eidechsen. Monatsber. Königl. Preuss. Akad. Wissensch. Berlin, 1869: 57-66 - get paper here
  • Peters, Wilhem Carl Hartwig 1869. Über neue Saurier (Chaunolæmus multicarinatus, Tropidolepisma richardi und Gymnodactylus steudneri) und Batrachier (Cyclohamphus fasciatus und Hyla gracilenta). Monatsber. königl. Akad. Wiss. Berlin. 1869 (November): 786-790 - get paper here
  • RAJABIZADEH, MAHDI; HIVA FAIZI, STEVEN C. ANDERSON, MOHAMMAD ZARRINTAB, ROMAN NAZAROV 2018. Taxonomic status of Tropiocolotes cf. steudneri with a description of a new species of Tropiocolotes (Reptilia: Squamata: Gekkonidae) in southern Iran. Zootaxa 4388 (2): 283–291 - get paper here
  • Rajabizadeh, Mehdi; Meisam Ghasemi, Hiwa Faizi, Nasrullah Rastegar-Pouyani 2010. New Data on Tropiocolotes cf. steudneri (Peters, 1869), (Sauria: Gekkonidae) in Southern Iran. Russ. J. Herpetol. 17 (3): 214-218 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • ROUNAQI, IMAN; ESKANDAR RASTEGAR-POUYANI, ALI GHOLAMIFARD, SAEED HOSSEINIAN 2017. Genetic and morphological variability among the populations assigned to the genus Tropiocolotes Peters, 1880 (Squamata: Gekkonidae) in south Iran. Zootaxa 4303 (2): 243–252 - get paper here
  • SAFAEI-MAHROO, BARBOD; HANYEH GHAFFARI, & STEVEN C. ANDERSON 2016. A new genus and species of gekkonid lizard (Squamata: Gekkota: Gekkonidae) from Hormozgan Province with a revised key to gekkonid genera of Iran. Zootaxa 4109 (4): 428–444 - get paper here
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Schlüter, U. 2004. Zwerggeckos (Tropiocolotes PETERS 1880) aus Nordafrika - Lebensräume, Verhalten, Pflege und Nachzucht. Draco 5 (18): 52-58 - get paper here
  • Shacham,B. & Shifman,S. 1998. The Tropiocolotes species in Israel is T. nattereri. Israel J. Zool. 44: 88
  • Shcherbak, N. N. [= Szczerbak], and M. L. Golubev 1996. Gecko fauna of the USSR and contiguous regions. SSAR [Russian original published in 1986]. [book review in Salamandra 36 (2): 138]
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Steindachner, F. 1901. Berichte der Comission für Oceanographische Forschungen. Expedition S. M. Schiff "Pola" in das Rothe Meer nördliche und südliche Hälfte. 1895/96 und 1897/98. Zoologische Ergebnisse. XVII. Bericht über die herpetologischen Aufsammlungen. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe, Wien, 69: 325-335 [1900]
  • Trape, J.F.; Trape, S. & Chirio, L. 2012. Lézards, crocodiles et tortues d'Afrique occidentale et du Sahara. IRD Orstom, 503 pp. - get paper here
 
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