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Uma thurmanae DERYCKE, GOTTSCHO, MULCAHY & DE QUEIROZ, 2020

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Higher TaxaPhrynosomatidae, Phrynosomatinae, Callisaurini; Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Mohawk Dunes Fringe-toed Lizard 
SynonymUma thurmanae DERYCKE, GOTTSCHO, MULCAHY & DE QUEIROZ 2020 
DistributionUSA (Arizona: Yuma County)

Type locality: the north end of the Mohawk Dunes (~2.2 km south of Interstate 8), Yuma County, Arizona, United States of America (32°41.698 N; 113° 48.935 W; WGS-84 datum).  
Reproduction 
TypesHolotype. USNM 590033, adult female; collected on 29 March 2015 by A. D. Gottscho at 17:28 hours. Deposited at the National Museum of Natural History, Smithsonian Institution, Washington, DC, USA.
Paratypes. USNM 590034 adult female, USNM 590035 adult male, USNM 590036 adult male, USNM 590037 juvenile female, USNM 590038 adult female, USNM 590039 juvenile female, USNM 590040 adult male, USNM 590041 adult male, USNM 590042 juvenile female, USNM 590043 adult male. Collected 25 and 26 March 2016 by A.D. Gottscho, J. Rabbers, and S. Harrington from Mohawk Dunes, Yuma County, AZ, USA. Deposited at the National Museum of Natural History, Washington, DC, USA. 
DiagnosisDiagnosis. Uma thurmanae is a morphologically cryptic species that doesn’t exhibit any known fixed morphological differences from other species in the complex, but it can be distinguished from those other species both by statistical differences in morphological characters (Table 5, Fig. 3) and in multivariate morphospace (Fig. 4). It can also be distinguished from all other members of the U. notata complex by nucleotide substitutions in three mtDNA loci (Table 8 in Derycke et al. 2020).

Coloration and pattern (in life). Dorsum with an ocellated pattern. The individual ocelli are roughly circular beige markings, each with a rusty central spot and radiating lines, surrounded by black. The lateral ocelli are aligned in rows so that their black borders form broken black longitudinal lines between the rows of ocelli. Underside of torso cream-colored and unblemished except for two black ventrolateral blotches slightly anterior to the midpoint between axilla and groin. Gular region marked by five black diagonal lines oriented anterolateral to posteromedial. Limbs lighter in coloration, consisting of a light-colored background with dark brown and gray spots. Eight black markings present ventrally on the tail, beginning 17.65 mm posterior to the cloaca as a small cluster of black dots in the center of the tail, and increasing in length and width posteriorly. In preservative, dorsal coloration is yellowed, with muted contrast between ocelli and background. 
Comment 
References
  • DERYCKE, ELIZABETH G.; ANDREW D. GOTTSCHO, ANDREW D. GOTTSCHO, DANIEL G. MULCAHY, KEVIN DE QUEIROZ 2020. A new cryptic species of fringe-toed lizards from southwestern Arizona with a revised taxonomy of the Uma notata species complex (Squamata: Phrynosomatidae). Zootaxa 4778 (1): 67–100 - get paper here
 
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