Vipera nikolskii VEDMEDERYA, GRUBANT & RUDAJEWA, 1986

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Higher Taxa	Viperidae, Viperinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Adder, Northern Viper G: Kreuzotter E: Nikolsky's Viper [nikolskii] G: Waldsteppen-Otter [nikolskii] Croatian: riđovka Russian: Обыкновенная гадюка
Synonym	Vipera nikolskii VEDMEDERYA, GRUBANT & RUDAJEWA 1986 Vipera berus var. prester — KOWATSCHEFF 1905 Vipera prester (LINNAEUS 1761) (fide KHALIKOV, pers. comm.) Vipera nikolskii — ENGELMANN et al 1993 Vipera berus nikolskii — JOGER et al. 1997 Vipera nikolskii — MCDIARMID, CAMPBELL & TOURÉ 1999: 407 Vipera berus nikolskii — MILTO & ZINENKO 2005 Vipera (Pelias) nikolskii — VENCHI & SINDACO 2006 Vipera berus nikolskii — VEDMEDERYA et al. 2009 Vipera berus nikolskii — ZINENKO et al. 2010 Pelias nikolskii — WALLACH et al. 2014: 544 Vipera nikolskii — BELIK 2014 Vipera berus nikolskii — FROLOVA & GAPONOV 2016 Pelias nikolskii — DUNAEV & ORLOVA 2017 Vipera berus nikolskii — MIZSEI et al. 2017 Vipera berus nikolskii — GENIEZ 2018: 293 Vipera nikolskii — ZAHER et al. 2019 Vipera berus nikolskii — FREITAS et al. 2020 Vipera berus nikolskii — JOGER et al. 2024
Distribution	Ukraine (vicinity of Kharkhov), Central and S Russia, Romania, Moldova Type locality: near the city of Kharkov, Ukraine
Reproduction	ovoviviparous (listed as viviparous by Blackburn 1994).
Types	Holotype: MNKhU (also as MNKNU) 14703.1
Diagnosis	Unfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 522 characters), are available for collaborators and contributors. Please contact us for details.
Comment	Synonymy: JOGER et al. (2007) found that bosniensis, nikolskii, barani, sachalinensis nested within various populations of V. berus. Type species: Vipera berus is the type species of the genus Pelias MERREM 1820. Pelias is the type genus of the subfamily Peliasinae REUSS 1933. Hybridization: Vipera berus appears to hybridize with V. nikolskii in Romania and Moldova (Zinenko et al. 2010).
Etymology	Apparently named after Alexander Mikhailovich Nikolsky (1858 – 1942), Russian and Ukrainian zoologist.
References	Belik V.P. 2014. To the history of research on the amphibian and reptile fauna in the Volgograd Region. Current Studies in Herpetology 14 (1/2): 3–13. Резюме - get paper here Cundall, D. 2009. Viper Fangs: Functional Limitations of Extreme Teeth. Physiological and Biochemical Zoology: Ecological and Evolutionary Approaches, 82(1), 63–79 - get paper here Dunaev E.A., Orlova V.F. 2017. Amphibians and reptiles of Russia. Atlas and determination. 2nd. ed. Moscow: Phyton XXI, 328 p Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp. Farooq, H., Uetz, P. 2021. Can we identify European snakes by color patterns? Salamandra 57 (4): 520–528 - get paper here Frolova E. N., Gaponov S. P. 2016. Morphology of Nikolsky’s viper in Voronezh and Lipetsk regions. PROCEEDINGS OF THE 6th CONGRESS OF THE A. M. NIKOLSKY HERPETOLOGICAL SOCIETY pp. 165-169 - get paper here Joger, Ulrich; Christophe Dufresnes & Sylvain ursenbacher 2024. Die Kreuzottergruppe (Vipera-berus-Komplex). Elaphe 2024 (3): 12-17 Joger,U., Lenk,P., Baran,I., Böhme,W., Ziegler,T., Heidrich,P. & M. Wink 1997. The phylogenetic position of Vipera barani and of Vipera nikolskii within the Vipera berus complex. In: Böhme,W. Bisschoff,W. & T. Ziegler (eds.) Herpetologia Bonnensis. Bonn: 185-194. - get paper here Kowatscheff, V. 1905. Beitraege zur Kenntnis der Reptilien und Amphibienfauna Bulgariens. Verh. Zool. Bot. Ges., Wien, IV: 31-32 McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp. Milto, K.D., Zinenko, O.I. 2005. Distribution and morphological variability of Vipera berus in eastern Europe. In: Herpetologia Petropolitana: Proceedings of 12th Ord. Gen. Meet. S.E.H., Ananyeva, N., Tsinenko, O., (Eds.). St. Petersburg – Moscow: 64 – 73 Vedmederja V I; Grubant V N; Rudayeva 1986. On the question of the name of the black viper in the forest steppes of the European part of the USSR [in Russian]. VESTNIK KHAR'KOVSKOGO UNIVERSITETA (No. 288): 83-85 - get paper here Vedmederya, Valeriy; Oleksandr Zinenko, and Andrei Barabanov 2009. An Annotated Type Catalogue of Amphibians and Reptiles in the Museum of Nature at V. N. Karazin Kharkiv National University (Kharkiv, Ukraine). Russ. J. Herpetol. 16 (3): 203-212 - get paper here Venchi, Alberto and Roberto Sindaco 2006. Annotated checklist of the reptiles of the Mediterranean countries, with keys to species identification. Part 2 -Snakes (Reptilia, Serpentes). Annali del Museo Civico di Storia Naturale "G. Doria", Genova, XCVIII: 259-364 Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Zaher H, Murphy RW, Arredondo JC, Graboski R, Machado-Filho PR, Mahlow K, et al. 2019. Large-scale molecular phylogeny, morphology, divergence-time estimation, and the fossil record of advanced caenophidian snakes (Squamata: Serpentes). PLoS ONE 14(5): e0216148 - get paper here Zinenko, Oleksandr; Ţurcanu, Vladimir; Strugariu, Alexandru 2010. Distribution and morphological variation of Vipera berus nikolskii Vedmederja, Grubant et Rudaeva, 1986 in Western Ukraine, The Republic of Moldova and Romania. Amphibia-Reptilia 31: 51-67 - get paper here
External links	National Center for Biotechnology Information Google images

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