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Xenosaurus manipulus NIETO-MONTES DE OCA, CASTRESANA-VILLANUEVA, CANSECO-MÁRQUEZ & CAMPBELL, 2022

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Higher TaxaXenosauridae, Diploglossa, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymXenosaurus manipulus NIETO-MONTES DE OCA, CASTRESANA-VILLANUEVA, CANSECO-MÁRQUEZ & CAMPBELL 2022: 41 
DistributionMéxico (Oaxaca)

Type locality: Vista Hermosa, km. 78 on Mexico Highway 175 Tuxtepec-Oaxaca, Municipality of Santiago Comaltepec, Oaxaca, México (17.63166667 N, 96.34250000 W), 1400 m elevation  
Reproduction 
TypesHolotype. MZFC-HE 9551, adult male, collected in a rock crevice in cloud forest on 15 October 1996 by Walter Schmidt Ballardo.
Paratypes. Twenty-two specimens, all from the Sierra de Jua ́rez, Oaxaca, Me ́xico: ‘‘Sierra de Jua ́rez’’ (no further data; MZFC-HE 2159 [three specimens]); Municipality of Santiago Comaltepec: Mexico Highway 175 Tuxtepec- Oaxaca, km. 88 (MZFC-HE 4510); La Esperanza (UTACV 22214); La Esperanza, 1550 m elevation (MZFC-HE 4511); 1 km NE La Esperanza (western slope of Cerro Redondo), 1736–1752 m elevation (MZFC-HE 9546–9547); same data as holotype (MZFC-HE 9548–9550); 3.4 mi. S Vista Hermosa (UTACV 8435); 3.8 mi. S Vista Hermosa (UTACV 8436); Ruta San Bernardo (approximately 3 km SW La Esperanza), 17.60711N, ????96.38464W, 1721 m (CNAR 17181–17182); northern face of Sierra Jua ́rez, 1.6 km S Vista Hermosa (UTACV 12116, 12118); northern face of Sierra Jua ́rez, 3.1 km S Vista Hermosa (UTACV 12120, 12124–12125); northern face of Sierra Jua ́rez, 4.8 km S Vista Hermosa (UTACV 12131); northern face of Sierra Jua ́rez, 6.8 km S Vista Hermosa (UTACV 12134).
Referred specimens. Six specimens, all from Oaxaca, Sierra de Jua ́rez, Municipality of Santiago Comaltepec: Campamento Vista Hermosa (KU 87437); Vista Hermosa (UCM 49320, 52482, 52487–52488); La Esperanza (UTACV 22213). 
Diagnosis 
CommentDistribution: see map in Nieto-Montes de Oca et al. 2022: 48 (Fig. 6). 
EtymologyNamed after the ancient Roman legion composed of either 60 or 120 men, and the origin of the modern word maniple. It is used in reference to the arrangement of the paravertebral tubercles of Xenosaurus manipulus, which resembles that of soldiers in the fighting position of the maniple. 
References
  • Nieto-Montes de Oca, Adrián; Nicté Castresana-Villanueva, Luis Canseco-Márquez & Jonathan A. Campbell 2022. A New Species of Xenosaurus (Squamata: Xenosauridae) from the Sierra de Juárez of Oaxaca, Mexico Herpetologica 78(1): 40–50. - get paper here
 
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