Philodryas erlandi LÖNNBERG, 1902

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Higher Taxa	Colubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Miranda Green Racer
Synonym	Philodryas Erlandi LÖNNBERG 1902 Philodryas ternetzii erlandi — MÜLLER 1928 Philodryas mattogrossensis — THOMAS 1976 (part.) Philodryas mattogrossensis — CEI 1993 (part.) Philodryas mattogrossensis — LOBO & SCROCCHI 1994 Philodryas mattogrossensis — MORO 1999 Philodryas mattogrossensis — ZAHER 1999 Philodryas mattogrossensis — ARZAMENDIA & GIRAUDO 2002 Philodryas mattogrossensis — KACOLIRIS et al. 2006 Philodryas mattogrossensis — SCHALK 2010 Philodryas erlandi — CACCIALI et al. 2016 Philodryas erlandi — KWET 2017 Philodryas erlandi — MELO-SAMPAIO et al. 2020
Distribution	Paraguay, SE Bolivia, N Argentina (Salta, Formosa, Chaco, Santiago del Estero) Type locality: Tarija, Bolivia
Reproduction	oviparous
Types	Syntypes: NRM 5097, BMNH 1946.1.8.43
Diagnosis	Diagnosis: Philodryas erlandi differs from all congeners except P. mattogrossensis by a dorsal colour that shifts from green on the anterior half of the body to brown on the posterior part. Philodryas erlandi is distinguished from P. mattogrossensis by the absence of a well-marked dark green or brown dorsal stripe restricted to the anterior one- third of the body (vs present in P. mattogrossensis), pres- ence of a dark line on the supralabials (vs absent or less evident in P. mattogrossensis), dorsal formula usually 19–19–15 (vs usually 21–19–15 in P. mattogrossensis), and first temporal elongated (vs. as wide as high in P. mattogrossensis).
Comment	Synonymy after CACCIALI et al. 2016. Distribution: see map in Cacciali et al. 2016: Fig. 11.
Etymology	Named in honor of Nils Erland Herbert Nordenskiöld (1877-1932), Swedish archeologist and anthropologist.
References	Arzamendia, V. & A. R. Giraudo 2002. Lista y distribución de los ofidios (Reptilia: Serpentes) de Santa Fe, Argentina. Cuadernos de Herpetología 16: 15–32 Cacciali, P., H. Cabral, V. L. Ferreira & G. Köhler 2016. Revision of Philodryas mattogrossensis with the revalidation of P. erlandi (Reptilia: Squamata: Dipsadidae). Salamandra 52 (4): 293-305 - get paper here Cei, J. M. 1993. Reptiles del noroeste, nordeste y este de la Argentina. Museo Regionale Sci. Naturale Torino, Monografie 14: 1-949 Kacoliris F.P.; Berkunsky I. & Williams J. 2006. Herpetofauna of Impenetrable, Argentinean Great Chaco. Phyllomedusa 5 (2): 149-158 - get paper here Kwet, Axel 2017. Neue Arten: Liste der im Jahr 2016 neu beschriebenen Reptilien. Terraria-Elaphe 2017 (3): 54-70 - get paper here Lobo, F. & G. Scrocchi 1994. Osteología craneal del género Philodryas (Serpentes: Colubridae). Cuadernos de Herpetología 8: 104–111 Lönnberg, E. 1902. On a collection of snakes from north-western Argentina and Bolivia containing new species. Ann. Mag. nat. Hist. (7) 10: 457-462 - get paper here Melo-Sampaio, P. R., Passos, P., Martins, A. R., Moura-leite, J. C., Morato, S. A., Venegas, P. J., Chávez G., Venâncio N.M. & De Souza, M. B. 2020. A phantom on the trees: integrative taxonomy supports a reappraisal of rear-fanged snakes classification (Dipsadidae: Philodryadini). Zoologischer Anzeiger 290: 19-39 [dated 2021 in print version, but 2020 published online] - get paper here Moro, S. 1999. Relaciones filogenéticas de Xenodontinae (Ser- pentes) neotropicales basadas en musculatura craneal. Cuadernos de Herpetología 13: 19–35 Müller, L. 1928. Herpetologische Mitteilungen. Zool. Anz. 77: 61-84 Schalk, Christopher M. 2010. Philodryas mattogrossensis: Diet. Herpetological Review 41 (3): 372 - get paper here Zaher, H. 1999. Hemipenial morphology of the South American xenodontine snakes, with a proposal for a monophyletic Xenodontinae and a reappraisal of colobrid hemipenes. Bull. Amer. Mus. Nat. Hist., 240: 1-168 - get paper here
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